After Lu, Zhu, Ahlberg, Qiao, Zhu, Zhao & Jia, 2016

<==o Sarcopterygii Romer, 1955 (flesh-finned osteichthyans; lihaseväiset kalat)
   |?- †Diabolepis speratus (Chang & Yu, 1984) Schultze, 1993; L.Dev. SEAs. [=Diabolichthyes (jr. synon.)] [Diabolichthyidae] [problematic, see Dipnoiformes]
   |?- †Langdenia campylognatha Janvier & Phuong, 1999; E.De. SEAs.
   |?- †Meemannia eos Zhu, Yu, Wang, Zhao & Jia, 2006; E.Dev. SEAs. [problematic, possibly basal actinopterygian (Lu et al., 2016)]
   |?- †Megamastax amblyodus Choo, Zhu, Zhao, Jia & Zhu, 2014; L.Sil. SEAs.
   |--+-- †Guiyu oneiros Zhu, Zhao, Jia, Lu, Qiao & Qu, 2009
   |  `--+-- †Psarolepis romeri Yu, 1998 sensu Lu et al., 2016; E.Dev. SEAs. [problematic, see Gnathostomata for other positions]
   |     `-- †Achoania jarvikii Zhu, Yu & Ahlberg, 2001
   `--+--o Coelacanthinimorpha sensu Lu et al., 2016 [Actinistia Cope, 1871] (coelacanths; latimeriansukuiset varsieväkalat)
      |  |-- †Bukkanodus jesseni Johanson, Long, Talenet, Janvier & Warren, 2007
      |  `--+-- †Onychodontida Andrews, 1973 [Onychodontiformes; Struniiformes]
      |     `-- Actinistia Cope, 1871 [Coelacanthinimorpha sensu Ahlberg, 1991, Cloutier & Ahlberg, 1996, Forey, 1998, Zhu, Yu & Janvier, 1999, and Clement, 2005]
      `--o Choanata [Rhipidistia] (choanates; sisäsieraimelliset)
         |-- †Styloichthys changae Zhu & Yu, 2002
         `--o Rhipidistia Cope, 1871 sensu Lu et al., 2016
            |-- Tetrapodomorpha Ahlberg, 1991 [Osteolepimorpha] [incl. Rhizodontida Andrews & Westoll, 1970; Osteolepida Boulenger, 1901] (Terrestrial vertebrate-like sarcopterygians; maaselkärankaismaiset lihaseväisetkalat)
            `--o Dipnomorpha Ahlberg, 1991 (lungfish-like sarcopterygians; keuhkokalamaiset lihaseväiset)
               |-- †Porolepiformes Jarvik, 1942
               `--o Dipnoiformes Cloutier, 1990 sensu Yu, 1998 (keuhkokalat; lungfishes)
                  |?- †Psarolepis romeri Yu, 1998 sensu Yu, 1998
                  `-- Dipnoiformes Cloutier, 1990 (keuhkokalat; lungfishes)

Tentative phylogenetic defenitions of sarcopterygian sub-clades:

Sarcopterygii = All osteichthyans closer to Homo sapiens (human) than Perca fluviatilis (common perch).

Coelacanthimorpha = All sarcopterygian closer to Latimeria chalumnae (coelacanth) tha either Protopterus annectens (west african lungfish) or Homo sapiens (human).

Choanata = All sarcopterygians closer to Protopterus annectens (west african lungfish) or Homo sapiens (human) than Latimeria chalumnae (coelacanth).

Rhipidistia = Last common ancestor of Protopterus annectens (west african lungfish) and Homo sapiens (human) and all it's decendants.

Family-level taxonomy of all non-tetrapod sarcopterygians.

Various proposed phylogenetic relationships of Sarcopterygian fishes (based on Cloutier & Ahlberg, 1996 and Lu & Zhu, 2009)

After Ahlberg, 1991, Cloutier & Ahlberg, 1996, Zhu, Yu & Janvier, 1999, and Zhu, Yu, Wang, Zhao & Jia, 2006

<==o Sarcopterygii
   |?- †Diabolepis
   |?- †Langdenia
   |-- †Meemannia
   |?- †Megamastax
   `--+?- †Psarolepis sensu Zhu, Yu, Wang, Zhao & Jia, 2006
      `--+-- †Onychodontida
         `--+-- Coelacanthinimorpha
            `--o Choanata
               |-- †Styloichthys
               `--+--o Tetrapodomorpha
                  |  |-- †Rhizodontida
                  |  `==o †Osteolepidida
                  |     `==o †Elpistostegalia
                  |        `-- Tetrapoda
                  `--o Dipnomorpha
                     |-- †Porolepiformes
                     `--o Dipnoiformes
                        |?- †Psarolepis sensu Yu, 1998
                        `-- Dipnoiformes

After Lu & Zhu, 2009

<==o Sarcopterygii
   |-- †Psarolepis
   `--+--+-- †Onychodontiformes
      |  `-- Actinistia
      `--+-- †Styloichthys
         `--+-- †Osteolepiformes
            `--+-- Dipnoi
               `-- †Porolepiformes

Reference(s):

  • Lu, J. & Zhu, M., 2009: An onychodont fish (Osteichthyes, Sarcopterygii) from the Early Devonian of China, and the evolution of the Onychodontiformes.
    –Proceedings of the Royal Society of London: Biological series: Vol. 277, #1, pp. 293-299 [doi: 10.1098/rspb.2009.0708]

After Zhu & Schultze, 1997 [from Campbell & Barwick, 2001]

<==o Sarcopterygii
   |?- †Meemannia
   |--+-- †Diabolepis
   |  `-- Dipnoi
   `--+-- †Youngolepis
      `--+-- †Powichthys
         `--+?- †Psarolepis
            `--+-- †Porolepiformes
               `--+-- Tetrapodomorpha
                  `--+-- †Onychodontida
                     `-- Coelacanthinimorpha

Reference(s):

  • Campbell, K. S. W. & Barwick, R. E., 2001: Diabolepis and its relationships to the Dipnoi.
    –Journal of Vertebrate Paleontology: Vol. 21, #2, pp. 227-241

After Schultze, 1987

<==o Sarcopterygii
   |-- Dipnoi
   `--+-- Actinistia
      `--+?- Onychodontiformes
         `--+-- †Powichthys & †Youngolepis
            `--+-- †Porolepiformes
               `--+-- †Osteolepiformes
                  `--+-- †Panderichthyidae
                     `-- Tetrapoda

After Panchen & Smithson, 1987

<==o Sarcopterygii
   |-- Actinistia
   `--+-- Dipnoi
      `--+--+-- †Powichthys
         |  `-- †Porolepiformes
         `--+-- †Youngolepis
            `--+-- †Osteolepiformes
               `-- Tetrapoda

After Rosen et al., 1981

<==o Sarcopterygii?
   |==“Osteolepiformes”
   `--+-- Actinistia
      |-- †Porolepiformes
      `--+-- Dipnoi
         `-- Tetrapoda

After Andrews, 1973

<==o Crossopterygii
   |--o Binostia
   |  |-- Coelacanthiformes
   |  `-- †Porolepiformes
   `--+-- Onychodontiformes
      `--o Quadrostia
         |-- †Rhizodontiformes
         `-- †Osteolepiformes

 

Reference(s):