After Ahlberg & Johanson, 1998, and Long, Young, Holland, Senden & Fitzgerald, 2006

Family-level taxonomy of non-tetrapod tetrapodomorphs
<==o Tetrapodomorpha Ahlberg, 1991 [Osteolepimorpha] (Terrestrial vertebrate-like sarcopterygians; maaselkärankaismaiset lihaseväiset)
   |-- †Tungsenia paradoxa Lu, Zhu, Long, Zhao, Senden, Jia & Qiao, 2012; E.Dev. SEAs.
   `--+-- †Kenichthys campbelli Chang & Zhu, 1993; E.Dev. SEAs.
      `--+-- †Rhizodontida Andrews & Westoll, 1970
         `--o Osteolepidida Boulenger, 1901 [Osteolepiformes Berg, 1937 [non Jarvik, 1942]]
            |--o †Osteolepididae
            |  |?- †Amadeodipterus kencampbelli Young & Schultze, 2005
            |  |?- †Luckeus abudda Young & Schultze, 2005
            |  |--o †Osteolepis Agassiz, 1835(?) [or Valenciennes, 1829] [Pleiopterus, Pliopterus, Triplopterus, Tripterus] M. Dev. NAs. Ant. M.-U. Dev. Eu.
            |  |  |-- †O. arenatus
            |  |  |-- †O. brevis
            |  |  |-- †O. macrolepidota (Sedgwick & Murchison, 1829)(?) [Osteolepis macrolepidotus; Thursius macrolepidotus (Sedgwick & Murchison, 1829)]
            |  |  |-- †O. major
            |  |  `-- †O. panderi
            |  `--+-- †Muranjilepis winterensis Young & Schultze, 2005
            |     `--o †Thursius Traquair, 1888
            |        |?- †T.? minor Jarvik, 1985
            |        |-- †T. estonicus Vorobyeva, 1977
            |        |-- †T. pholidotus Traquair, 1888
            |        |-- †T. talsiensis Traquair, 1888
            |        |-- †T. truncatus Traquair, 1888
            |        |-- †T. wudingensis Fan, 1992
            |        `-- †T. macrolepidotes (Sedgwick & Murchison, 1829)
            |?-o †Thysanolepidae [= Megalichthyidae??]
            |  |-- †Thysanolepis Vorobyeva, 1977
            |  `-- †Vorobjevaia dolonodon Young, Long & Ritchie, 1992; L.Dev. Ant.
            `--+--o †Megalichthyidae Long, 1985(?)
               |  |?- †Megistolepis klementzi (Obruchev, 1955) Vorobjeva, 1977
               |  |?- †Megapomus markovskyi Vorobyeva, 1977
               |  |-- †Mahalalepis resima Young, Long & Ritchie, 1992; L.Dev. Aust.
               |  |?- †Sengoerichthys ottoman Janvier, Clement & Cloutier, 2007
               |  |-- †Ectosteorhachis nitidus Cope, 1880?
               |  `--+-- †Cladarosymblema narrienense Fox et al., 1995
               |     `--o †Megalichthys
               |        |-- †M. hibberti Agassiz, 1843
               |        |-- †M. laticeps Traquair, 1884
               |        |?- †M. agassizianus Lohest, 1889
               |        |?- †M. coccolepis Young, 1870
               |        |?- †M. intermedius Woodward, 1891
               |        `?- †M. macropomus Cope, 1892
               `--+-- †Medoevia lata Lebedev, 1995
                  `--+-- †Gogonasus andrewsae Long, 1985; L.Dev. Aust.
                     |--o †Canowindridae Young, Long & Ritchie, 1992
                     |  |?- †Owensia chooi Holland, 2009
                     |  |-- †Beelarongia patrichae Long, 1987
                     |  `--+-- †Koharalepis jarviki Young, Long & Ritchie, 1992 [Koharolepis?? (lapsus calami?)] L.Dev. Ant.
                     |     `-- †Canowindra grossi Thomson, 1973
                     `--+--o †Gyroptychius Jarvik, 1948
                        |  |-- †G.? australis Young & Gorter, 1981
                        |  |-- †G. agassizi Jarvik, 1948
                        |  |-- †G. milleri Jarvik, 1948
                        |  |-- †G. groenlandicus Jarvik, 1950
                        |  `-- †G. dolichotatus Jarvik, 1985
                        `--+-- †Tristichopteridae Cope, 1889
                           `--o †Elpistostegalia Camp & Allison, 1961 [Panderichthyida Vorobyeva, 1981]
                              |-- †Tinirau clackae Swartz, 2012, M.Dev. WNAm.
                              |?- †Parapanderichthys stolbovi (Vorobyeva, 1960) Vorobyeva, 1992 [Panderichthys stolbovi Vorobyeva, 1960]
                              `--+--o †Platycephalichthys Vorobyeva, 1959 [previously treated as tristichopterid]
                                 |  |-- †P. skuenicus Vorobyeva, 1962
                                 |  |-- †P. bischoffi Vorobyeva, 1959
                                 |  `-- †P. rohoni Vorobyeva, 1962
                                 `--+-- †Panderichthys rhombolepis Gross, 1941 [incl.? Panderichthys bystrowi Gross, 1941] [Panderichthyidae Vorobyeva, 1968]
                                    `--+-- †Elpistostege watsoni Westoll, 1938 [Elpistostegidae]
                                       |-- †Howittichthys warrenae Long & Holland 2008
                                       |-- †Livoniana multidentata Ahlberg, Luksevic & Mark-Kurik, 2000
                                       |-- †Tiktaalik roseae Daeschler, Shubin & Jenkins, 2006
                                       `-- Stegocephali Laurin, 1998a [Tetrapoda Haworth, 1825 sensu Goodrich 1930, and Laurin & Reisz, 1997] (tetrapods = vertebrates with four recognizible limbs; maaselkärankaiset)


Osteolepida incertae sedis [taxa not treated by Ahlberg & Johanson, 1998]:

    †Bogdanovia orientalis Obrucheva 1955; L.Dev. CAs. [has been treated as Coelacanthinimorph sarcopterygian]
    †Canningius groenlandicus Säve-Söderbergh, 1937
    †Chrysolepis [non Chrysolepis (chinquapins), a genus of fabaceaen plants (Fabaceae)]
    †GeiserolepisLatvius grewingki (Gross, 1933)
    †Lohsania utahensis Vaughn, 1962
    †Megadonichthys kurikae Vorobyeva, 1962
    †Platyethmoidia antarctica Young, Long & Ritchie, 1992; L.Dev. Ant.
    †Shirolepis ananjevi Vorobeva, 1977
    †Sterropterygion brandei Thomson, 1972
    †Thaumatolepis edelsteini Obruchev, 1941

Reference(s):

  • Ahlberg, P. E., Clack, J. A. & Luksevics, E. 1996: Rapib braincase evolution between Panderichthyes and the earliest tetrapods.
    –Nature: vol. 381, 2 May, pp. 61-64
  • Ahlberg, P. E., Luksevics, E. & Mark-Kurik, E., 2000: A near-tetrapod from the Baltic Middle Devonian.
    –Palaeontology: Vol. 43, #3, pp. 533-548
  • Ahlberg, P. E. & Milner, A. R. 1994: The origin and early diversification of tetrapods.
    –Nature: vol. 368, 7 April, pp. 507-514.
  • Ahlberg, P. E. & Johanson, Z. 1998: Osteolepiforms and the ancestry of tetrapods.
    –Nature: Vol. 395, 22 October, pp. 792-794
  • Blom, H., Clack, J. A., Ahlberg, P. E. & Friedman, M., 2007: Devonian vertebrates from East Greenland: a review of faunal composition and distribution.
    –Geodiversitas: Vol. 29, #1, pp. 119-141
  • Carroll, R. L., 1988: Vertebrate paleontology and evolution.
    –W. H. Freeman and company, New York, 1988, 698
  • Carroll, R. L., 1988: Appendix. 594-648
    in Carroll, R. L., 1988: Vertebrate paleontology and evolution.
    –W. H. Freeman and company, New York, 1988, 698
  • Carroll, R. L. 1996: Revealing the patterns of macroevolution.
    –Nature: vol. 381, 2 May, pp. 19-20.
  • Daeschler, E. B., Shubin, N. H. & Jenkins Jr, F. A., 2006: A Devonian tetrapod-like fish and the evolution of the tetrapod body plan.
    –Nature: Vol. 440, pp. 757-763 [doi: 10.1038/nature04639]
  • Frickhinger, K. A., 1995: Fossil Atlas – Fishes.
    –Mergus – Publishers for Natural History and Pet Books, Hans A.Baensch, Malle, Germany, 1995, 1-1088
  • Lebedev, O. A., 1995: Morphology of a new osteolepidid fish from Russia.
    –Bulletin du Muséum national d'Histoire naturelle, Paris, 4e sér.: Vol 17, Section C, n° 1-4, pp. 287-341
  • Long, J. A., 1995: The Rise of Fishes: 500 million years of evolution.
    –Johns Hopkins University Press, Baltimore & London, pp. 1-223
  • Long, J. A., Young, G. C., Holland, T., Senden, T. J. & Fitzgerald, E. M. G., 2006: An exceptional Devonian fish from Australia sheds light on tetrapod origins.
    †Nature: Vol. 444, #7116, pp. 199-202 [doi:10.1038/nature05243]
  • Lu, J., Zhu, M., Long, J. A., Zhao, W., Senden, T. J., Jia, L. & Qiao, T., 2012: The earliest known stem-tetrapod from the Lower Devonian of China.
    –Nature Communications: Vol. 3, pp. 1160 [doi: 10.1038/ncomms2170]
  • Johanson, Z. & Ahlberg, P. E., 1998: A complete primitive rhizodont from Australia.
    –Nature: Vol. 394, 6 August, pp. 569-573
  • Swartz, B., 2012: A Marine Stem-Tetrapod from the Devonian of Western North America.
    –Plos One: Vol. 7, #3, pp. e33683(1-11) [doi: 10.1371/journal.pone.0033683]
  • Young, G. C., Long, J. A. & Ritchie, A., 1992: Crossopterygian fishes from the Devonian of Antarctica: systematics, relationships and biogeographic signifcance.
    –Records of the Australian Museum: Supplement 14, pp. 1-77 [doi: 10.3853/j.0812-7387.14.1992.90]
  • Young, G. C. & Schultze, H.-P., 2005: New osteichthyans (bony fishes) from the Devonian of Central Australia.
    –Mitteilungen aus dem Museum für Naturkunde in Berlin – Geowissenschaftliche Reihe: Vol. 8, #1, pp. 13-35 [from abstract]