After Clement & Long, 2010, and Lloyd, Wang & Brusatte, 2012 (based on datasets from Schultze & Marshall, 1993, Schultze, 2001 and 2004); with untreated taxa from Carroll, 1988, Cloutier & Ahlberg, 1996
Note: Lloyd et al. don't give any supernominal names for nodes in their phylogenetic tree. Names which I have given to various nodes are those which has been previously used for Dipnoi. This is one of the aspects that clearly speak for comprehensive phylogeny-based taxonomic treatment for Dipnoi.
<==o Dipnoiformes Cloutier, 1990 [Dipnoi Müller, 1845] (lungfishes; keuhkokalat) |--o †Powichthys Jessen, 1975 [Powichthyidae Jessen, 1980] L.Dev. NNA. Spits. | |-- †P. thorsteinssoni Jessen, 1975 | `-- †P. spitsbergensis Clément & Janvier, 2004 `--+-- †Youngolepis preacursor Chang & Yu, 1981 `--o Dipnoi Müller, 1845 [Dipneusti] |-- †Diabolepis speratus (Chang & Yu, 1984) Schultze, 1993 [=Diabolichthyes, preoccupied] [Diabolepididae Schultze, 1993] `--+--o ?†Dipnorhynchidae Berg, 1940 | |-- †Westollrhynchus lehmanni (Westoll, 1949) Schultze, 2001 [Dipnorhynchus lehmanni Westoll, 1949; Speonesydrion lehmanni (Westoll, 1949) Campbell & Barwick, 1985] | `--+--o †Ichnomylax Long, Campbell & Barwick, 1994 | | |-- †I. kurnai Long, Campbell & Barwick, 1994 | | `-- †I. karatajae Reisz, Krupina & Smith, 2004 | `==o †Dipnorhynchus Jaekel, 1927; L.Dev. Eu., M.Dev. Aust. | |-- †D. suessmilchi (Etheridge) Hills, 1941, Thomson & Campbell 1971 [Ganorhynchus sussmilchi Etheridge] [type] | `--+-- †“Chirodipterus” onawwayensis Schultze, 1982 | `--+-- †D. cathlesae Campbell & Barwick, 1999 | `-- †D. kurikae Campbell & Barwick, 1985 `--+-- †“Dipnorhynchus” kiandrensis Campbell & Barwick 1982 `--+-- †Uranolophus wyomingensis L. Dev. NA. [Uranolophina: Uranolophidae Miles, 1977] `--+-- †Speonesydrion iani Campbell & Barwick, 1983 `--+-- †Jarvikia arctica Lehman, 1959 `--+--+--o ?†Dipteridae Owen, 1846 | | |-- †Barwickia downunda Long, 1992a | | `--o †Dipterus Sedgwick & Murchison, 1828 [Catopterus, Eoctenodus, Paradipterus, Polyphractus] L.-U. Dev. Eu. M.-U. Dev. NA. U. Dev. NAs. Aus. | | |?- †D. oervigi Gross, 1964 | | `-- †D. valenciennesi Sedgwick & Murchison, 1828 [incl. Palaeospondylus gunni Traquair [Palaeospondyliformes: Palaeospondylidae:], a larval form] | `--o ?†Chirodipteridae Campbell & Barwick, 1990 | |--+-- †Dipterus cf. valenciennesi | | `-- †Chiropterus rhenanus Mörs, 1991 | `--+-- †Adololopas moyasmithi Campbell & Barwick, 1998 | `--o †Chirodipterus Gross, 1933; U. Dev. Eu. Aus. | |-- †C. australis Miles, 1977 | |?- †C. liangchengi Song & Chang, 1991 | |?- †C. potteri Kemp, 2000 | `-- †C. wildungensis (Gross, 1933) [non Säve-Söderbergh, 1952] `--+-- †Pillararhynchus longi Campbell & Barwick, 1990 [Chirodipteridae: Pillararhynchinae Long, 1992] `--+--+-- †Sorbitorhynchus deleaskitus Wang, Drapala, Barwick & Campbell, in Campbell & Barwick, 1990 [Chirodipteridae: Pillararhynchinae Long, 1992] | `-- †Melanognathus canadensis Jarvik, 1967; L. Dev. NNA. `--+-- †Palaeodaphus insignis (Gervais) [Palaedaphus; ?Archaeonectes; ?Heliodus; ?Paleodahus]; U.Dev. Eu. NAm. `--+-- †Sunwapta grandiceps Thomson, 1967 `--+-- †Gogodipterus paddyensis (Miles, 1977) Long, 1992b [Chirodipterus paddyensis Miles, 1977] `--+--o †Stomiahykidae Bernacsek, 1977 | |-- †Stomiahykus thlaodus Bernacsek, 1977; M.Dev. NA. | `-- †Amadeodipterus kencampbelli Young & Schultze, 2005; M.Dev. CAust. `--+-- †Tarachomylax oepiki Barwick, Campbell & Mark-Kurik, 1997 `--+-- †Iowadipterus halli Schultze, 1992 `--+--o †Rhinodipterus Gross, 1959; M.-U. Dev. Eu. [Rhinodipteridae Campbell & Barwick, 1990] | |-- †R. secans Gross, 1959 | |-- †R. ulrichi Ørvig, 1961 | |-- †R. stolbovi Krupina, 1995 | `-- †R. kimberleyensis Clement, 2012 `--+--o †Phaneropleuridae Huxley, 1861 | |-- †Oervigia nordica Lehman, 1959; U.Dev. Gr. | `-- †Phaneropleuron andersoni U. Dev. Eu. ?Gr. `--+?- †Harajicadipterus youngi Clement, 2009; M.Dev CAust. |--o ?†Holodipterida Moy-Thomas & Miles, 1971 | |-- †Orlovichthys limnatis Lehman, 1959 [non? Krupina, 1980]; U. Dev. EEu. | `--+--+--+-- †Scaumenacia curta (Whiteaves 1881) Traquair, 1893 [Canadipterus Jaekel, 1927; Canadiptarus (lapsus calami)] U. Dev. NA. ?Eu. | | | `-- †Pentlandia macroptera (Traquair, 1888) [Dipterus macroptera Traquair, 1888] M. Dev. Eu. | | `--+-- †Howidipterus donnae Long, 1992a | | `-- †Delatitia breviceps (Woodward, 1906) Long & Campbell, 1985 | `--+--o †Fleurantiidae Berg, 1940 | | `-- †Fleurantia denticulata Graham Smith & Westoll 1937; U.Dev. NA. | `--o ?†Holodontoidei Schultze, 1992 | |--o †Rhynchodipteridae Moy-Thomas, 1939 [non? Berg, 1940] | | |--o †Griphognathus Gross, 1959 | | | |-- †G. minutidens Gross, 1959 | | | |-- †G. sculpta Schultze, 1969(?) | | | `-- †G. whitei Miles, 1977 | | `--+-- †Rhynchodipterus elginensis Säve-Söderbergh, 1937 | | `--+-- †Jessenia concentrica Otto & Bardenheuer, 1996 [non Jessenia, a synonym of Oenocarpus (a Brazilian palmtree)] | | `--o †Soederberghia Lehman, 1959; U. Dev. Gr. Aus. NA. | | |-- †S. groenlandica Lehman, 1959 | | `-- †S. simpsoni Ahlberg, Johanson & Daeschler, 2001 | `--o †Holodontidae Gorizdro-Kulczycka, 1950 sensu Clement & Long, 2010b [HolodipteridaMoy-Thomas & Miles, 1971: Holodontoidei Schultze, 1992] | |--+-- †Archaeonectes pertusus Meyer, 1858 | | `-- †Robinsondipterus longi (Campbell & Barwick, 1991) Long, 2010 [Holodipterus longi Campbell & Barwick, 1991; Holodipterus (Holodipterus) longi Pridmore, Campbell & Barwick, 1994; Holodipterus (Robinsondipterus) longi] | `--+--o †Holodipterus White & Moy-Thomas, 1940 [Archaeotylus; Holodus; Holodipterus (Holodipterus) White & Moy-Thomas; Pridmore, Campbell & Barwick, 1994] U.Dev. EEu. Aust. | | |-- †H. kiprijanovae (Pander) [type, suppressed?] | | `-- †H. gogoensis Miles, 1977 [Holodipterus (Holodipterus) gogoensis Pridmore, Campbell & Barwick, 1994] | `--+-- †Xeradipterus hatcheri Clement & Long, 2010b; U.Dev. Aust. | `--+-- †“Holodipterus” (Holodipteroides) elderae Pridmore, Campbell & Barwick, 1994 | `-- †Asthenorhynchus meemannae (Pridmore, Campbell & Barwick, 1994) Long, 2010 [Holodipterus (Asthenorhynchus) meemannae Pridmore, Campbell & Barwick, 1994] `--+--+-- †Adelargo schultzeiJohanson & Ritchie, 2000; U.Dev. Aust. | `--o †Andreyeichthys Krupina, 1987 [non? Krupina & Reisz, 1999] SWNAm. | |?- †A. sp. | `-- †A. epitomus Krupina, 1987 `--+-- †“Holodipterus” sanctacrucensis Gorizdro-Kulczycka, 1950 `-- “Post-Devonian dipnoans” [Ceratodiformes sensu lato]
Untreated Dipnoi:
†Cathlorhynchus trismodipterus Campbell, Barwick & Senden, 2009
†Ganorhynchus Traquair, 1873; M.-U.Dev. Eu. NAm.
†G. beecheri (Newberry)
†G. rigauxi (Dutertr, 1931)
†G. woodwardi Traquair, 1878
†G. splendens Gross, 1937
†Erikia jarviki Chang & Wang, 1995
†Iranorhynchus seyedemamii Janvier & Martin, 1978
†Osteoplax erosus M'Coy, 1848; Penn. WEu.
†Pinnalongus saxoni Newman & den Blaauwen, 2007; M.Dev. WEu.
†Sinodipterus beibei Qiao " Zhu, 2009; M.Dev. EAs.
†Dipteridae Owen, 1846
†Conchodus jerofejewi (Pander, 1858) McCoy, 1848 [Creatodus lateralis: Eichwald, 1846; Cheirodus jerofejewi Pander, 1858; Cheirodus lateralis Eichwald, 1860; Conchodus jerofejewi (Pander, 1858) Krupina, 2004] M.-U. Dev. Eu. U. Dev. NA.
†Dipteroides Jaekel, 1926 (or 1935); U.Dev. Eu.
Alternative phylogeinies of Dipnoiformes:
After Clement & Long, 2010 (fig. 10b)
<==o Dipnoiformes
|-- †Diabolepis
|-- †Tarachomylax
`--+-- †Jessenia
|-- †Stomiahykus
|--+-- †Uranolophus
| `--+-- †Dipnorhynchus
| `--+-- †Ichnomylax
| `-- †Speonesydrion
`--+-- †Gogodipterus
`--+--+-- †Melanognathus
| `--+--+-- †Griphognathus
| | `-- †Robinsondipterus
| `--+-- †Holodipterus gogoensis
| `--+-- †Xeradipterus
| `--+-- †“Holodipterus” (Holodipteroides) elderae
| `-- †Asthenorhynchus
`--+--+-- †Chirodipterus australis
| `-- †Sorbitorhynchus
`--+-- †Dipterus
`--+-- †Adololopas
`--+-- †Pillararhynchus
`--+-- †Orlovichthys
`--+--+-- †Soederberghia
| `-- †Conchopoma
`--+-- †Andreyevichthys
`--+--+-- †Scaumenacia
| `--+-- †Fleurantia
| `--+-- †Barwickia
| `-- †Howidipterus
`--+-- †Ctenodus
|-- †Sagenodus
`--+-- †Gnathorhiza
`-- Neoceratodus
|
After Qiao & Zhu, 2009
<==+-- †Youngolepis
`--o Dipnoi
|-- †Diabolepis
`--+--+-- †Westollrhynchus
| `--+-- †Ichnomylax
| `--+-- †Dipnorhynchus sussmilchi
| `-- †Speonesydrion
`--+-- †Uranolophus
`--+-- †Stomiahykus
`--+--+-- †Iowadipterus
| `-- †Jessenia
`--+-- †Tarachomylax
|-- †Melanognathus
|-- †Gogodipterus
`--+-- †Dipterus
`--+--+-- †Adololopas
| `--+-- †“Chirodipterus” liangchangi
| `--+-- †“Chirodipterus” australis
| `--+-- †Sorbitorhynchus
| `-- †Pillararhynchus
`--+-- †Sinodipterus
`--+--+-- †Rhinodipterus
| `-- †Chirodipterus wildungensis
|--+-- †Holodipterus
| `--+-- †Soederberghia
| `-- †Griphognathus
`--+-- †Adelargo
|-- †Fleurantia
|--+-- †Howidipterus
| `-- †Barwickia
`--+-- †Scaumenacia
`--+-- †Andreyevichthys
`-- †Orlovichthys
Reference(s):
|
After Clement, 2009 (Fig. 4b)<==o Dipnoiformes |-- †Diabolepis |--+-- †Jessenia | `--+-- †Uranolophus | `--+-- †Stomiahykus | |-- †Dipnorhynchus sussmilchi | `--+-- †Speonesydrion | `-- †Ichnomylax `--+--+-- †Tarachomylax | `-- †Melanognathus `--+-- †Dipterus |--o “Chirodipterid clade” | |-- †Adololopas | `--+-- †Sorbitorhynchus | |-- †Pillararhynchus | `--+-- †Gogodipterus | `-- †Chirodipterus australis `--+-- †Harajicadipterus |--o “Holodontid clade” | |--+-- †Holodipterus gogoensis | | `-- †Holodipterus (Asthenorhynchus) meemannae | `--+-- †Holodipterus (Robinsondipterus) longi | `-- †Griphognathus whitei `--+-- †Orlovichthys `--+--+-- †Conchopoma | `-- †Soederberghia `--+-- †Andreyevichthys `--+--+-- †Scaumenacia | `--+-- †Fleurantia | `--+-- †Howidipterus | `-- †Barwickia `--+-- †Sagenodus |-- †Ctenodus `--+-- †Gnathorhiza `-- Neoceratodus Reference(s):
|
After Friedman, M., 2007 (Fig. 19., 20.)
<==o Dipnoiformes
|-- †Powichthys
`--+-- †Youngolepis
`--+-- †Diabolepis
`--+--+-- †Dipnorhynchus sussmilchi
| `-- †Dipnorhynchus kurikae
`--+-- †Uranolophus
|-- †Stomiahykus
`--+?- †Melanognathus
`--+--+-- †“Chirodipterus” australis
| `-- Gogodipterus
`--+-- †Pillararhynchus
`--+-- †Sorbitorhynchus
`--+-- †Dipterus valenciennesi
`--+?- †“Chirodipterus” liangchengi
`--+-- †Orlovichthys
`--+-- †“Chirodipterus” wildungensis
`--+-- †Holodipterus gogoensis
`--+-- †“Griphognathus” whitei
`--+-- †Griphognathus minutidens
`-- †Soederberghia
Reference(s):
|
After Schultze, 2001
<==o Dipnoiformes
|-- †Psarolepis
|-- †Diabolepis
`--+--+-- †Uranolophus
| `-- †Melanognathus
`--+-- †Westollrhynchus sp.
`--+--+--+-- †D. kurikae
| | `-- †D. kiandrensis
| `--+-- †“Speonesydrion” iani
| `-- †D. suessmilchi
`--+-- †Stomiahykus
`--+-- †Jessenia
`--+-- †Iowadipterus
`--+--+-- †Pillararhynchus
| `-- †Sorbitorhynchus
`--+-- †Tarachomylax
`--+-- †Dipterus
`--+-- †Adololopas
`--+-- †Rhinodipterus
`--+--+-- †Gogodipterus
| `-- †Chirodipterus
`--+--+-- †Holodipterus
| `-- †Adelargo
`--+--+-- †Howidipterus
| `--+-- †Barwickia
| `--+-- †Fleurantia
| `--+-- †Griphognathus
| `--+-- †Rhynchodipterus
| `--+-- †Soederberghia
| `-- †Jarvikia
`--+-- †Scaumenacia
`--+-- †Pentlandia
`--+-- †Oervigia
`--+-- †Andreyevichthys
`--+-- †Phaneropleuron
`-- †Orlovichthys
Reference(s):
|
After Carroll, 1988, and Cloutier & Ahlberg, 1996<==o Dipnoiformes |--o †Powichthyidae | `-- †Powichthys `--+-- †Youngolepis `--o Dipnoi |-- †Diabolepis `--+--o †Uranolophina | `--o †Uranolophidae | `-- †Uranolophus `--o Dipnorhynchina |?-o †Holodontidae | |-- †Adelargo | `-- †Holodipterus |--o †Rhynchodipteridae | `-- †Rhinodipterus |--o †Dipnorhynchidae | |-- †Dipnorhynchus | |-- †Ganorhynchus | |-- †Griphognathus | `-- †Holodipterus |--o †Sagenodontidae | |?- †Proceratodus | |-- †Megapleuron | |-- †Sagenodus | `-- †Straitonia `--o Speonesydrionina |?- †Osteoplax |?- †Palaeophichthys |--o †Dipteridae | |?- †Palaedaphus | |?- †Stomiahykus | |-- †Chirodipterus | |-- †Conchodus | |-- †Dipteroides | |-- †Grossipterus | |-- †Pentlandia | `-- †Dipterus `--+?-o †Fleurantiidae | `-- †Fleurantia `--+--o †Phaneropleuridae | |-- †Jarvikia | |-- †Nielsenia | |-- †Oervigia | |-- †Phaneropleuron | |-- †Scaumenacia | `-- †Soederberghia |--o †Ctenodontidae | |-- †Ctenodus | `-- †Tranodis `--+--o †Conchopomidae | `-- †Conchopoma `--+--o †Uronemidae | `-- Uronemus `--o Ceratodiformes |-- †Arganodus |-- †Gosfordia |-- †Microceratodus |-- †Paraceratodus |--o †Asiatoceratodontidae | `-- †Asiatoceratodus |--o †Ptychoceratodontidae | `-- †Ptychoceratodus |--o †Ceratodontidae | `-- †Ceratodus |--o Neoceratodontidae | |-- †Mioceratodus | `-- Neoceratodus `--o Lepidosirenidae |-- †Gnathorhiza |-- Lepidosiren `-- Protopterus Reference(s):
|
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