After Long, 1995, Cloutier & Ahlberg, 1996, Forey, 1998, and Clement, 2005
Family-level taxonomy of Coelacanthinimorpha<==o Coelacanthinimorpha [Actinistia Cope, 1871] (coelacanths or Latimeria-like sarcopterygians; latimeriamaiset varsieväkalat) |?- †Shoshonia arctopteryx Friedman, Coates & Anderson, 2007; M.-L.Dev. NAm. |--o †Miguashaidae Schultze, 1993 [paraphyletic?] | |-- †Gavinia syntrips Long, 1999; M.Dev. Aus. | `--o †Miguashaia Schultze, 1973; M.Dev. Eu. NAm. | |-- †M. grossi Forey et al., 2000; M.Dev. Eu. | `-- †M. bureaui Schultze, 1973; M.Dev. NAm. `--+?-†Eoactinistia foreyi Johanson, Long, Talent, Janvier & Warren, 2006: E.Dev. SEAust. |--o †Diplocercidae Stensiö, 1922 | |-- †Dictyonosteus Stensiö, 1918; L.Dev. Spits. | `--o †Diplocercides Stensiö, 1922 [Holoptychius von Koenen, 1895; Coelacanthus Woodward, 1898; Nesides Stensiö, 1937] L.Dev. NEEu. | |-- †D. kayseri (von Koenen, 1895) Stensiö, 1922 [Holoptychius kayseri von Koenen, 1895; Coelacanthus? kayseri (von Koenen) Woodward, 1898; Nesides schmidti Stensiö, 1937] L.Dev. Eu. | |-- †D. davisi (Moy-Thomas, 1937) [Coelacanthus abdenensis Traquair, 1903; Rhabdoderma davisi Moy-Thomas, 1937; Rhabdoderma(?) abdenense Moy-Thomas, 1937] L.Dev. WEu. | |-- †D. jaekeli Stensiö, 1922, L.Dev. CEu. | `-- †D. heiligenstockiensis (Jessen, 1966) [Nesides? heiligenstockiensis Jessen, 1966] L.Dev. CEu. `--+?-o †Euporosteus Jaeckel, 1927 M.Dev. Eu. SEAs. [Euporosteidae] | |-- †E. eifeliensis Jaeckel, 1927 [E. eifelianus] | `-- †E. yunnanensis Zhu, Yu, Lu, Qiao, Zhao & Jia, 2012 |?- †Chagrinia enodis Schaeffer, 1962; L.Dev. NAm. |?-o †Hadronectoroidei Lund & Lund, 1984 | |?- †Lochmocercus aciculiodontus Lund & Lund, 1984; E.Carb. (Miss.) NAm. | `--o †Hadronectoridae Lund & Lund, 1984 | |-- †Allenypterus montanus Melton, 1969; E.Carb. (Miss.) NAm. | |-- †Hadronector donbairdi Lund & Lund, 1984; E.Carb. (Miss.) NAm. | |-- †Holopterygius nudus Jessen, 1973; L.Dev. CEu. | `-- †Polyosteorhynchus simplex Lund & Lund, 1984; E.Carb. (Miss.) NAm. `--+?=o †“Rhabdodermatidae” Berg, 1958 [paraphyletic] | |-- †Cardiosuctor populosus Lund & Lund, 1984; E.Carb. (Miss.) NAm. | |-- †Synaptotylus newelli (Hibbard, 1933) Echols, 1963 [Coelacanthus newelli Hibbard, 1933; Coelacanthus arcuatus Hibbard, 1933; Rhabdoderma? newelli (Hibbard) Forey, 1998] L.Carb. (Penn.) NAm. | `--o †Rhabdoderma Reis, 1888 [Taxonomic history] Miss. NAf. Miss. Penn. Eu. EEu. NAm. | |-- †R.? alderingi Moy-Thomas, 1937 | |-- †R. elegans (Newberry, 1856) [Taxonomic history] | |-- †R. ardrossense Moy-Thomas, 1937 | |-- †R. exiguum (Eastman, 1902) [Taxonomic history] | |-- †R. huxleyi (Traquair, 1881) [Taxonomic history] | `-- †R. tingleyense (Davis, 1884) [Taxonomic history] `--+--o †Sasseniidae Forey, 1998 | |-- †Spermatodus pustulosus Cope, 1894; E.Perm. NAm. | `--o †Sassenia Stensiö, 1921; L.Trias. Spits. Gr. | |?- †S.? guttata (Woodward, 1912) [Coelacanthus guttata Woodward, 1912] | |-- †S. tuberculata Stensiö, 1921 | `-- †S. groenlandica Forey, 1998 `--+--o †Laugiidae Stensiö, 1932 | |-- †Belemnocerca prolata Wendruff & Wilson, 2013; E.Trias. WNAm. | |-- †Laugia groenlandica Stensiö, 1932; E.Trias. Gr. | |?- †Trachymetopon liassicum Aldinger, 1951; E.Jur. CEu. | `--o †Coccoderma Quenstedt, 1858 emend. Reis, 1888 [Kokkoderma (sic) Quenstedt, 1858] [non Coccoderma Möllendorff, 1901, a pulmonate gastropod genus (Enidae)] L.Jur. CEu. | |-- †C.? bavaricum Reis, 1888 [a species of Undina? (Forey, 1998)] | |-- †C.? gigas Reis, 1888 | |-- †C. harlemensis (Winkler, 1871) [Coelacanthus harlemensis Winkler, 1871] | |-- †C.? substriolatum (Huxley, 1866) Reis, 1888 | `-- †C. suevicum Quenstedt, 1858 [Coccoderma nudum Reis, 1888] `--+?-o †Whiteiidae Schultze, 1993 [treated as coelacanthiniforms by Forey (1998) and Clement (2005)] | |-- †Guizhoucoelacanthus guanlingensis Liu et al., 2006 sensu Geng, Zhu & Jin, 2009 | `--+-- †Piveteauia madagascariensis Lehman, 1952; E.Trias. Mad. | `--o †Whiteia Moy-Thomas, 1935 E.Trias. Mad. Gr. ?E. Trias. NA. | |-- †W. woodwardi Moy-Thomas, 1935 [Coelacanthus evolutus Beltan, 1980] E.Trias. Mad. | |-- †W. africanus (Broom, 1905 [Coelacanthus africanus Broom, 1905] E.Trias. SAf. | |-- †W. tuberculata Moy-Thomas, 1935; E.Trias. Mad. | `-- †W. sp. (Gardiner, 1966); E.Trias. WNAm. `-- Coelacanthiformes Huxley, 1861 sensu Clement, 2005 (latimeriat; coelacanths)
Coelacanthinimorpha incertae sedis:
†Changxingia Wang & Liu, 1981; L.Perm. CAs. †Changxingia aspratilis Wang & Liu, 1981 †C. weii Jin, 1997 †Chouhuichthys majiashanensis Tong, Zhou, Erwin, Zuo & Zhao, 2006; L.Trias. EAs. [a coelacanthid (Tong, et al., 2006)] †Coelocanthopsis curta Traquair, 1901; L.Carb. (Miss.) Eu. EEu. †Cualabaea L.Jur. WAf. †Graphiurichthys callopterus (Kner, 1866) White & Moy-Thomas, 1937 [Graphiurus Kner, 1866] L.Trias. Eu. †Hainbergia granulata Schweizer, 1966; M.Trias. CEu. †Heptanema paradoxum Bellotti, 1857 [non Stensiö, 1932] M.-L.Trias. Eu. †Moenkopia wellesi Schaeffer & Gregory, 1961; M.Trias. WNAm. †Mylacanthus Stensiö, 1921 E.Trias. Spits. †Mylacanthus lobatus Stensiö, 1921 E.Trias. Spits. †Mylacanthus spinosus Stensiö, 1921 E.Trias. Spits. †Rhipis de Saint-Seine, 1950; L.Cret. CAf. [scales] †Rhipis moorseli de Saint-Seine, 1950 †Rhipis tuberculatus Casier, 1965 †Rhipis sp. [L.Cret. CAf.(Casier, 1965)] †Scleracanthus asper Stensiö, 1921; E.Trias. Spits. †Sinocoelacanthus fengshanesis Liu, 1964; E.Trias. EAs. †Youngichthys xinhuainsis Wang & Liu, 1981; L.Perm. CAs.
Alternative phylogenies of Coelacanthomorpha:
After Cavin, Mennecart, Obrist, Costeur & Furrer, 2017<==o Coelacanthinimorpha |-- †Miguashaia `--+-- †Diplocercides `--+-- †Lochmocercus |--+-- †Rebellatrix | `-- †Hadronector `--+--+-- †Allenypterus | |-- †Holopterygius | `-- †Euporosteus `--+-- †Polyosteorhynchus `--+-- †Cardiosuctor `--+-- †Rhabdoderma `--+--+-- †Spermatodus pustulosus | `-- †Sassenia `--+--+-- †Piveteauia | `--+-- †Laugia | `-- †Coccoderma `--o Coelacanthiformes(?) |-- †Coelacanthus `--+-- †Guizhoucoelacanthus `--+-- †Whiteia `--+--+-- †Wimania | `-- †Axelia `--+-- †Dobrogeria `--+-- †Yunnancoelacanthus `--+-- †Luopingcoelacanthus `--o Latimerioidei |--o †Mawsoniidae | |-- †Diplurus | `--+-- †Chinlea | |-- †Parnaibaia | |-- †Lualabaea | `--+-- †Mawsonia | `-- †Axelrodichthys `--o Latimeridae |-- †Gambergia `--+--+-- †Megalocoelacanthus | `-- †Libys `--+--+-- †Ticinepomis | `-- †Foreyia `--+-- †Holophagus |-- †Undina |-- †Indocoelacanthus `--+-- †Macropoma `--+-- †Swenzia `-- Latimeria Reference(s):
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After Wen et al., 2013 (fig. 8a)<==o Osteichthyes |-- †Allenypterus `--+-- †Holopterygius `--+-- †Hadronector |-- †Lochmocercus |--+-- †Diplocercides | `--+-- †Miguashaia | `--+-- Porolepiformes [+ Dipnoi??!?] | `-- Actinopterygii `--o Coelacanthiformes (??) |-- †Polyosteorhynchus `--+-- †Cardiosuctor `--+-- †Rhabdoderma `--+--+-- †Sassenia | `-- †Spermatodus `--o Coelacanthoidei |--+-- †Coccoderma | `-- †Laugia `--+-- †Coelacanthus `--+-- †Piveteauia `--+-- †Guizhoucoelacanthus `--+-- †Whiteia `--+-- †Luopingcoelacanthus `--+-- †Yunnancoelacanthus `--o Latimerioidei |--o †Mawsoniidae | |-- †Diplurus | `--+-- †Chinlea | `--+-- †Axelrodichthys | `-- †Mawsonia `--o Latimeriidae |-- †Granbergia |-- †Libys |--+-- †Undina | `-- †Holophagus `--+-- †Macropoma `--+-- †Swenzia `-- Latimeria Refence(s):
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After Dutel, Maisey, Schwimmer, Janvier, Herbin & Clément, 2012<==o Coelacanthinimorpha |-- †Miguashaia `--+--+-- †Diplocercides | `-- †Euporosteus `--+-- †Hadronector |-- †Lochmocercus `--+--+-- †Allenypterus | `-- †Holopterygius `--+-- †Polyosteorhynchus `--+-- †Cardiosuctor `--+-- †Rhabdoderma `--+-- †Spermatodus pustulosus |-- †Sassenia |--+-- †Laugia | `-- †Coccoderma `--o Coelacanthiformes(?) |--+-- †Piveteauia | `--+-- †Whiteia | `--+-- †Guizhoucoelacanthus | |-- †Axelia | `-- †Wimania |-- †Coelacanthus `--+-- †Rebellatrix `--+-- †Granbergia `--o Latimerioidei |--o †Mawsoniidae | |-- †Diplurus | `--+-- †Chinlea | |-- †Parnaibaia | |-- †Lualabaea | `--+-- †Mawsonia | `-- †Axelrodichthys `--o Latimeridae |-- †Ticinepomis `--+-- †Holophagus |-- †Undina |-- †Indocoelacanthus |--+-- †Megalocoelacanthus | `-- †Libys `--+-- †Macropoma `--+-- †Swenzia `-- Latimeria Reference(s):
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After Wendruff & Wilson, 2012<==o Coelacanthinimorpha |-- †Miguashaia `--+-- †Diplocercides `--+-- †Hadronector `--+--+-- †Allenypterus | `-- †Holopterygius `--+-- †Lochmocercus `--+-- †Polyosteorhynchus `--+--+-- †Cardiosuctor | `-- †Rhabdoderma `--+-- †Sassenia `--+--+-- †Coelacanthus | `--+-- †Coccoderma | `-- †Laugia `--+-- †Spermatodus `--+-- †Whiteia `--+-- †Rebellatrix `--o Latimerioidei |-- †Granbergia |--o †Mawsoniidae | |-- †Chinlea | `--+-- †Parnaibaia | `--+-- †Axelrodichthys | `-- †Mawsonia `--+--+-- †Diplurus | `-- †Libys `--o Latimeriidae |--+-- †Undina | `-- †Holophagus `--+-- †Macropoma `--+-- †Swenzia `-- Latimeria Reference:
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After Geng, Zhu & Jin, 2009 (fig. 4a)<==o Coelacanthinimorpha |-- †Miguashaia `--+-- †Diplocercides `--+-- †Hadronector |-- †Lochmocercus `--+--+-- †Allenypterus | `-- †Holopterygius `--+-- †Polyosteorhynchus `--+-- †Cardiosuctor `--+-- †Rhabdoderma `--+--o †Sasseniidae | |-- †Sassenia | `-- †Spermatodus `--+--o †Laugiidae | |-- †Coccoderma | `-- †Laugia `--o Coelacanthiniformes |--o †Whiteiidae | |-- †Guizhoucoelacanthus | `--+-- †Piveteauia | `-- †Whiteia `--+-- †Coelacanthus `--o Latimerioidei |--o †Mawsoniidae | |-- †Granbergia | `--+-- †Chinlea | `--+-- †Axelrodichthys | `-- †Mawsonia `--+--+-- †Diplurus | `-- †Libys `--o Latimeriidae |--+-- †Undina | `-- †Holophagus `--+-- †Macropoma `--+-- †Swenzia `-- Latimeria Reference:
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After Forey, 1998<==o Actinistia |-- †Miguashaia `--+-- †Diplocercides `--+-- †Hadronector |-- †Lochmocercus `--+-- †Allenypterus `--+-- †Polyosteorhynchus `--+-- †Cardiosuctor `--+-- †Rhabdoderma `--+--o †Sasseniidae | |-- †Sassenia | `-- †Spermatodus `--+--o †Laugiidae | |-- †Coccoderma | `-- †Laugia `--o Coelacanthiniformes |-- †Coelacanthus `--+-- †Whiteia `--o Latimerioidei |--o †Mawsoniidae | |-- †Granbergia | `--+-- †Libys | `--+-- Diplurus | `--+-- †Chinlea | `--+-- †Axelrodichthys | `-- †Mawsonia `--o Latimeriidae |--+-- †Undina | `-- †Holophagus `--+-- †Macropoma `-- Latimeria Reference(s):
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