After Prothero & Schoch, 1989, McKenna & Bell, 1997 and Thewissen, Williams & Hussain, 2001

<==o Helaletidae Osborn, 1892 [paraphyletic?]
   |?- †Hyrachyus asiaticus
   |?- †Jhagirilophus chorgalensis Thewissen, Williams & Hussain, 2001
   |?- †Triplopus kalakotensis
   |-- †Helaletes nanus (Marsh, 1871a) [Veragromovia, Chasmotheroides]
   |-- †Desmatotherium intermedius (Osborn, Scott & Spier, 1878) [Helaletes intermedius]
   |-- †Selenaletes scopaeus
   |-- †Dilophodon minusculus Scott, 1883 [Heteraletes]
   |-- †Haagella
   |-- †Plesiocolopirus hancocki
   `--o †Colodon
      |?- †C. cingulatus
      |-- †C. kayi
      |-- †C. woodi
      `-- †C. occidentalis

Reference(s):

  • Carroll, R. L., 1988: Vertebrate paleontology and evolution.
    –W. H. Freeman and company, New York, 1988, 698.
  • Carroll, R. L., 1988: Appendix. 594-648.
    in Carroll, R. L., 1988: Vertebrate paleontology and evolution.
    –W. H. Freeman and company, New York, 1988, 698.
  • Gazin, C. L., 1976: Mammalian faunal zones of the Bridger Middle Eocene.
    –Smithsonian Contributions to Paleobiology: No. 26, pp. iv-25
  • McKenna, M. C. & Bell, S. K., (eds.) 1997: Classification of mammals – above the species level.
    –Columbia University Press, New York, 1997, xii-631
  • Prothero, D. R. & Schoch, R. M., 1989: Classification of the Perissodactyla. 530-537.
    in Prothero, D. R. & Schoch, R. M., (eds.) 1989: The Evolution of Perissodactyls.
    –Oxford University Press, New York, New York & Oxford, England, 1989, ix-537
  • Thewissen, J. G. M., Williams, E. M. & Hussain, S. T., 2001: Eocene mammals faunas from Northern Indo-Pakistan.
    –Journal of Vertebrate Paleontology: Vol. 21, #2, pp. 347-366