Mikko's Phylogeny Archive Main Groups of Non-Therian Mammals   Holotheria

After Rougier, Novacek, McKenna & Wible, 2001

<==o Holotheria Wible et al., 1995 sensu Luo, Cifelli & Kielan-Jaworowska, 2001 [Theria sensu lato] ("korkeammat nisäkkäät")
   |?- †Chronoperates Fox, Youzwyshyn & Krause, 1992 [Chronoperatidae Fox, Youzwyshyn & Krause, 1992]
   `--o Trechnotheria McKenna, 1975 [Yangotheria Chow & Rich, 1982]
      |?- †Liaotherium gracile Zhou, Cheng & Wang, 1991
      |--+-- †Gobiconodontidae Chow & Rich, 1984 sensu Luo, Kielan-Jaworowska & Cifelli, 2002
      |  `-- †Triconodontidae Marsh, 1887 sensu Rougier, Novacek, McKenna & Wible, 2001
      `--+--+-- †Aploconodon comoensis Simpson, 1925
         |  `--+--o †Amphiodontidae Simpson, 1925 [Amphiodontoidea Prothero, 1981]
         |     |  |-- †Amphiodon superstes Simpson, 1925
         |     |  |-- †Gobiotheriodon infinitus (Trofimov, 1980) [Gobiodon Trofimov, 1980]
         |     |  `--+-- †Manchurodon simplicidens Yabe & Shikama, 1938
         |     |     `-- †Nakunodon paikasiensis Yadagiri, 1985
         |     `--+-- †Hakusanodon
         |        `-- †Comodon gidleyi (Simpson, 1925) Kretzoi & Kretzoi, 2000 [Phascolodon Simpson, 1925; Phascolotheridium Cifelli RL & Dykes TD, 2001]
         `--+-- †Jeholodens jenkinsi Ji, Luo & Ji, 1999
            |-- †Amphilestes broderipii Owen, 1845
            |-- †Phascolotherium bucklandi Broderip, 1828 [Didelphys Broderip, 1828; Thylacotherium Valenciennes, 1838]
            `--o Symmetrodonta Simpson, 1925
               |?- †Casamiquelia Bonaparte, 1990
               |?- †Thereuodon Sigogneau-Russell, 1989
               |?- †Atlasodon Sigogneau-Russell, 1991
               |-- †Eurylambda aequicrurius
               `--+--+-- †Zhangheotherium quinquecuspidens Hu, Wang, Luo & Li, 1997 [transfered to Spalacotheroidea by Li & Luo, 2006]
                  |  `--+-- †Peralestes longirostris
                  |     `-- †Spalacotheroidea Prothero, 1981 [Spalacotheriida Prothero, 1981, incl. Quirogatheria Bonaparte, 1992]
                  `--o Mammalia Linnaeus, 1758 sensu stricto [conclusion after Luo, Cifelli & Kielan-Jaworowska, 2001]
                     |?- †Tribactonodon bonfieldi Sigogneau-Russell, Hooker, Ensom, 2001
                     |--+?-o †Shuontherium Chow & Rich, 1982 [Yinotheria Chow & Rich, 1982: Shuotherida Chow & Rich, 1982: Shuotheriidae Chow & Rich, 1982]
                     |  |  |-- †S. dongi Chow & Rich, 1982
                     |  |  |-- †S. shilongi Wang, Clemens, Hu & Li, 1998
                     |  |  `-- †S. kermacki Sigogneau-Russell, 1998
                     |  `-- Australophenida Luo, Cifelli & Kielan-Jaworowska, 2001 [incl. Monotremata and Ausktribosphenida]
                     `-- Cladotheria McKenna, 1975 sensu Luo, Cifelli & Kielan-Jaworowska, 2001

Note:

This tree is based on alternative phylogeny of Holotherians presented by Rougier, Novacek, McKenna & Wible, 2001. While doing this tree I made some presumptions not expressed in their original tree (Fig. 6).

• I presume that they chose Amphiodon to present all other Amphiodontids as well.
• I presume that Eurylambda presents Symmetrodonta
• I presume that Amphilestes presents Cladotheria
• I also turned their node 21 to get Cladotheria as most derived taxa in this tree.

I have also included other taxa not present in their analysis, and where indoubt, I included sistergroups from Kielan-Jaworowska, Cifelli & Luo, 2002.

Reference(s):

Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698

Carroll, R. L., 1988: Appendix. 594-648
in Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698

Cifelli, R. L., Wible, J. R. & Jenkins, F. A. jr., 1998: Triconodont mammals from the Cloverly Formation (Lower Cretaceous), Montana and Wyoming.
–Journal of Vertebrate Paleontology: Vol. 18, #1, pp. 237-241

Dykes, T., 2002: Personal correspondence.

Dykes, T., 2001: Spalacotheriidae, Tinodontidae and other derived symmetrodonts.
–iNet: Mesozoic mammals – an internet directory: http://home.arcor.de/ktdykes/symmetro.htm

Kielan-Jaworowska, Z., Cifelli, R. L. & Luo, Z.-X., 2002: Dentition and relationships of the Jurassic mammal Shuotherium.
–Acta Palaeontologica Polonica: Vol. 47, #3, pp. 479-486

Luo, Z. 1994: Sister-group relationships of mammals and transformations of diagnostic mammalian characters. 98-128.
in Fraser, N. C. & Sues, H-D. (eds.) 1994: In the Shadow of the Dinosaurs.
–Cambridge University Press, New York, 1994, x-435

Luo, Z.-X., Cifelli, R. L. & Kielan-Jaworowska, Z., 2001: Dual origin of tribosphenic mammals.
–Nature: Vol. 409, 4 January, pp. 53-57

Luo, Z.-X., Kielan-Jaworowska, Z. & Cifelli, R. L., 2002: In quest for a phylogeny of Mesozoic mammals.
–Acta Palaeontologica Polonica: Vol. 47, #1, pp. 1-78

McKenna, M. C. & Bell, S. K., (eds.) 1997: Classification of mammals – above the species level.
–Columbia University Press, New York, 1997, xii-631

Rougier, G. W., Novacek, M. J., McKenna, M. C. & Wible, J. R., 2001: Gobiconodonts from the Early Cretaceous of Oshih (Ashile), Mongolia.
–American Museum Novitates: #3348, pp. 1-30

Wang, Y., Clemens, W. M., Hu, Y. & Li, C., 1998: A probable pseudo-tribosphenic upper molar from the Late Jurassic of China and the early radiation of the Holotheria.
–Journal of Vertebrate Paleontology: vol. 18, #4, pp. 777-787