After Luo, Cifelli, & Kielan-Jaworowska, 2001 and Luo, Kielan-Jaworowska & Cifelli, 2002

Alternative phylogeny after McKenna & Bell, 1997

Alternative phylogeny after Wang, Clemens, Hu & Li, 1998

Alternative phylogeny after Rougier, Wible, Beck & Apesteguía, 2012

<==o Cladotheria McKenna, 1975 sensu McKenna & Bell, 1997, and Luo, Cifelli, & Kielan-Jaworowska, 2001
   |?- †Butlerigale Kühne, 1968 [possibly pathological Dryolestes leiriensis]
   |--o †Dryolestoidea Butler, 1939 [Dryolestiformes Kinman, 1994; incl. Pantotheria, Marsh, 1880]
   |  `-- †Dryolestida Prothero, 1981
   `--o Zatheria McKenna, 1975
      |?- †Arguitherium Dashzeveg, 1994 [Arguitheriidae Dashzeveg, 1994]
      |?- †Arguimus Dashzeveg, 1979 [Arguimuridae Dashzeveg, 1979]
      |?- †Abelodon Brunet et al., 1991
      |?-o †Nanolestes T. Martin, 2002
      |  |-- †N. drescherae T. Martin, 2002
      |  `-- †N. krusati T. Martin, 2002
      |?- †Minimus richardfoxi Sigogneau-Russell, 1999
      |?- †Magnimus ensomi Sigogneau-Russell, 1999
      |?- †Afriquiamus nessovi Sigogneau-Russell, 1999
      |-- †Vincelestes neuquenianus Bonaparte, 1986 [Vincelestidae Bonaparte, 1986]
      `--+-- Boreosphenida Luo, Cifelli, & Kielan-Jaworowska, 2001
         `--+--o †Amphitherium de Blainville, 1838 [taxonomic history] [Amphitheriidae Owen, 1846; Amphitheriida Prothero, 1981]
            |  |?- †A. oweni
            |  |-- †A. rixoni Butler & Clemens, 2001
            |  `-- †A. prevostii de Blainville, 1838
            `-- †Peramura McKenna, 1975

After McKenna & Bell, 1997

<==o Cladotheria
   |?- †Butlerigale
   |?-o †Ausktribosphenidae
   |  |?- †Bishops
   |  `-- †Ausktribosphenos
   |--o †Dryolestoidea
   |  |-- †Dryolestida
   |  `--o †Amphitheriida
   |     `--o †Amphitheriidae
   |        `-- †Amphitherium
   `--o Zatheria
      |?- †Arguitherium
      |?- †Arguimus
      |?- †Nanolestes
      |?- †Vincelestes
      |-- †Peramura
      `-- Tribosphenida

Reference(s):

  • McKenna, M. C. & Bell, S. K., (eds.) 1997: Classification of mammals – above the species level.
    –Columbia University Press, New York, 1997, xii-631

After Wang, Clemens, Hu & Li, 1998

<==o Cladotheria
   |?- †Butlerigale
   |--o †Dryolestoidea
   |  `-- †Dryolestida
   `--o Amphitheriida
      |-- †Amphitherium
      `--o Zatheria
         |?- †Arguitherium
         |?- †Arguimus
         |?- †Nanolestes
         |?- †Vincelestes
         |-- †Peramura
         `-- Tribosphenida

Reference(s):

  • Wang, Y., Clemens, W. M., Hu, Y. & Li, C., 1998: A probable pseudo-tribosphenic upper molar from the Late Jurassic of China and the early radiation of the Holotheria.
    –Journal of Vertebrate Paleontology: vol. 18, #4, pp. 777-787

After Rougier, Wible, Beck & Apesteguía, 2012

<==o Cladotheria
   |-- †Dryolestidae
   `--+-- †Paurodontidae
      `--+-- †Meridiolestida
         `-- Boreosphenida

Reference(s):

  • Rougier, G. W., Wible, J. R., Beck, R. M. D. & Apesteguía, S., 2012: The Miocene mammal Necrolestes demonstrates the survival of a Mesozoic nontherian lineage into the late Cenozoic of South America.
    –Proceedings of the National Academy of Sciences: in press. [doi: 10.1073/pnas.1212997109]

Reference(s):