After Meylan & Ganko, 1988 [from Meylan & Ganko, 1997, and Meylan, 2001] and Anquetin, Barrett, Jones, Moore-Fay & Evans, 2009 (fig. 3d)

<==o Chelonia Macartney, 1802 (turtles; kilpikonnat) [alternative phylogenetic position after Rieppel & deBraga, 1996]
   |-- †Pappochelys rosinae Schoch & Sues, 2015; M.Trias. CEu.
   `--+-- †Odontochelys semitestacea Li, Wu, Rieppel, Wang & Zhao, 2008; L.Trias. SAs.
      `--+-- †Australochelys africanus Gaffney & Kitching, 1994 [Australochelidae sensu Lee, 1997]
         `--o Rhaptochelydia Gaffney & Kitching, 1994 [Testudines Linnaeus, 1758 [non Batsch, 1788] sensu Lee, 1996]
            |-- †Chinlechelys tenertesta Joyce, Lucas, Scheyer, Heckert & Hunt, 2009; L.Trias. WNAm.
            |-- †Proganochelys quenstedtii Baur, 1887; L.Trias. CEu. [Chelytherium; Psammochelys; Stegochelys; Triassochelys] [Proganochelydia: Proganochelyidae]
            |?-o †Proterochersidae Nopcsa, 1923a sensu Szczygielski & Sulej, 2016
            |  |-- †Keuperotesta limendorsa Szczygielski & Sulej, 2016
            |  `--o †Proterochersis Fraas, 1913 [Murrhardtia Karl & Tichy, 2000]
            |     |-- †P. robusta Fraas, 1913 [Proterochersis intermedia Fraas, 1913; Murrhardtia staeschei Karl & Tichy, 2000]
            |     `-- †P. porebensis Szczygielski & Sulej, 2016
            `--+?- †Palaeochersis talampayensis Rougier, de la Fuente & Arcucci, 1995; L.Trias. SAm.
               |?- †Condorchelys antiqua Sterli, 2008; M.Jur., SSAm.
               |?-+-- †Eileanchelys waldmanni Anquetin, Barrett, Jones, Moore-Fay & Evans, 2009; M.Jur. WEu.
               |  `-- †Hackerochelys romani
               `--o Casichelyidia Gaffney, 1975 [Testudinata Klein, in Behn, 1760 (non Linnaeus, 1758) sensu Lee, 1997; Testudines Treviranus, 1802] (recent turtles; nykykilpikonnat)
                  |-- Pleurodira Cope, 1864 (fide Gaffney & Maylan, 1988) [Pan-Pleurodira Joyce, Parham & Gauthier, 2004] (side-necked turtles; sivutaivuttajat)
                  `-- Cryptodira Dumeril & Bibron, 1835 [non Cope, 1871] [Pan-Cryptodira Joyce, Parham & Gauthier, 2004; Cryptodiramorpha Lee, 1995 sensu Parham & Hutchison, 2003] (cryptodires; pystytaivuttajat)

Testudinata incertae sedis:

Chitracephalus dumonii Dollo, 1884
        †Neusticemys neuquina Fernandez & de la Fuente,, 1988
        †Scutemys appalanata Wiman, 1930 [Scutemys tecta Wiman, 1930]
        †Tacuarembemys kusterae Perea, Soto, Sterli, Mesa, Toriño, Roland, & Da Silva, 2014
    †Chelycarapookidae Warren, 1969
        †Chelycarapookus arcuatus Warren, 1969

Alternative phylogenies of basal turtles:

After Szczygielski & Sulej, 2016

<==o Testudinata
   |-- †Odontochelys
   `--+--o †Proterochersidae
      |  |-- †Keuperotesta
      |  `-- †Proterochersis
      `--+-- †Proganochelys
         `--+-- †Australochelys
            `--+-- †Palaeochersis
               `--+-- †Kayentachelys
                  `-- “Post-Early Jurassic turtles”

Reference(s):

  • Szczygielski, T. & Sulej, T., 2016: Revision of the Triassic European turtles Proterochersis and Murrhardtia (Reptilia, Testudinata, Proterochersidae), with the description of new taxa from Poland and Germany.
    –Zoological Journal of the Linnean Society: Vol. 177, #2, pp. 395-427 [doi: 10.1111/zoj.12374]

After Rabi, Zhou, Wings, Ge & Joyce, 2013

<==o Testudinata
   |-- †Odontochelys
   `--+-- †Proganochelys
      `--+--+-- †Australochelys
         |  `-- †Palaeochersis
         `--+-- †Proterochersis
            `--+-- †Indochelys
               |-- †Condorchelys
               |-- †Kayentachelys
               `--+-- †Heckerochelys
                  `--+-- †Eileanchelys
                     `--+-- †Meiolaniformes
                        `--+-- †Changyuchelys
                           `--+-- †Paracryptodira
                              `--+-- †Xinjiangchelys
                                 `--+-- †Basilochelys
                                    `--+-- †Sinemys
                                       `--+-- †Ordosemys
                                          `--+-- †Dracochelys
                                             `--+-- Judithemys
                                                |-- Hangaiemys
                                                `--o Testudines
                                                   |-- Pleurodira
                                                   `-- Cryptodira s.s.

Reference(s):

  • Rabi, M., Zhou, C.-F., Wings, O., Ge, S. & Joyce, W., 2013: A new xinjiangchelyid turtle from the Middle Jurassic of Xinjiang, China and the evolution of the basipterygoid process in Mesozoic turtles.
    –BMC Evolutionary Biology: Vol. 13, #1, pp. (1-29) [doi: 10.1186/1471-2148-13-203]

After Joyce, Lucas, Scheyer, Heckert & Hunt, 2009 (fig. 3)

<==o  Testudinata
   |-- †Chinlechelys
   |-- †Proganochelys
   `--+-- †Palaeochersis
      |-- †Australochelys
      `--+-- †Proterochersis
         `--+-- †Kayentachelys
            `--+-- †Mongolochelys
               `--o Testudines
                  |-- Pleurodira
                  `-- Cryptodira

After Anquetin, Barrett, Jones, Moore-Fay & Evans, 2009 (fig. 3d)

<==o Testudinata
   |-- †Proganochelys
   `--+-- †Palaeochersis
      `--+-- †Australochelys
         `--+-- †Proterochersis
            `--+-- †Kayentachelys
               `--+?- †Condorchelys antiqua
                  |--+-- †Eileanchelys
                  |  `-- †Heckerochelys
                  `--+--+-- †Mongolochelys
                     |  `-- †Meiolania
                     `--+-- †Kallokibotion
                        `--o Testudines
                           |-- Pleurodira
                           `-- Cryptodira

After Li, Wu, Rieppel, Wang & Zhao, 2008

<==o Testudinata
   |-- †Odontochelys
   `--+-- †Proganochelys
      `--+-- †Palaeochersis
         |-- †Proterochersis
         `--o Testudines
            |-- Pleurodira
            `-- Cryptodira

After Sterli, 2008

<==o Testudinata
   |-- †Proganochelys [Proganchelys Sterli, 2008 (lapsus calami)]
   `--+--+-- †Palaeochersis
      |  `-- †Australochelys
      `--+-- †Proterochersis
         `--+-- †Kayentachelys
            |-- †Condorchelys
            |-- †Indochelys
            `--+-- †Heckerochelys
               `--+--+-- †Mongolochelys
                  |  `-- †Meiolania
                  `--+-- †Kallokibotion
                     `--o Testudines
                        |-- Pleurodira
                        `-- Cryptodira

After Rougier, de la Fuente & Arcucci, 1995

<==o Testudines
   |-- †Proganochelys
   `--+--o †Australochelidae
      |  |-- †Palaeochersis
      |  `-- †Australochelys
      `--o Rhaptochelydia
         |-- †Proterochersis
         `--o Casichelyidia
            |-- Pleurodira
            `--o Cryptodira
               |-- †Kayentachelys
               `-- Selmacryptodira

After Gaffney & Kitching, 1994(?)

<==o Chelonia
   |--o †Australochelidae
   |  `-- †Australochelys
   `--o Rhaptochelydia
      |--o †Proganochelydia
      |  `--o †Proganochelyidae
      |     `-- †Proganochelys
      `--o Casichelyidia
         |-- Pleurodira
         `-- Cryptodira

 

Reference(s):