After Maylan & Ganko, 1997, Lapparent de Broin, 2000, and Gaffney, Tong & Buffetaut, 2006, with species from the “Paleobiology Database”
<==o Pleurodira Cope, 1864 (fide Gaffney & Maylan, 1988) [Pan-Pleurodira Joyce, Parham & Gauthier, 2004] (side-necked turtles; sivutaivuttajat) |--o †Proterochersidae Nopcsa, 1928 [Minipleurodira Gaffney, Tong & Buffetaut, 2006] | |?- †Saurischiocomes keuperinus Kuhn 1939 | `--o †Proterochersis Fraas, 1913 [Chelytherium obscurum Meyer, 1863 (nomen oblitum), Murrhardtia Karl & Tichy 2000] | |-- †P. robusta Fraas, 1913 [Proterochersis robustum Fraas, 1913; Murrhardtia staeschei Karl and Tichy 2000] | `-- †P. intermedia Fraas 1913 `--o Megapleurodira Gaffney, Tong & Buffetaut, 2006 |?-o †Dortokidae Lapparent de Broin & Murelaga, 1996 | |-- †“Muehlbachia nopcsai” Vremir & Codrea, 2009 [nomen nudum/nomen invalidum] L.Cret. EEu. | |--+-- †Eodortoka morellana Pérez-García, Gasulla & Ortega, 2014; E.Cret. SWEu. | | `-- †Dortoka vasconica Lapparent de Broin & Murelaga, 1996; L.Cret. WEu. | `-- †Ronella botanica Gheerbrant, Codrea, Hosu, Sen, Guernet, Lapparent de Broin & Riverline, 1999; Paleoc. EEu. |--o †Platychelira Gaffney, Tong & Buffetaut, 2006 | `--o †Platychelyidae Brän, 1965 sensu Gaffney, Tong & Buffetaut, 2006 | |-- †Platychelys oberndorferi Wagner, 1853 | `--o †Notoemys Cattoi & Freiberg, 1961 [incl. †Caribemys de la Fuente & Iturralde-Vinent, 2001] | |-- †N. zapatocaensis Rueda & Gaffney, 2005 | |-- †N. oxfordiensis (de la Fuente & Iturralde-Vinent, 2001) [†Caribemys oxfordiensis de la Fuente & Iturralde-Vinent, 2001]; L. Jur. Car. | `-- †N. laticentralis Cattoi & Freiberg, 1961 `--o Eupleurodira Gaffney & Maylan, 1988 sensu Gaffney, Tong & Buffetaut, 2006 |--o Cheloides Gray, 1825 sensu Gaffney, Tong & Buffetaut, 2006 | `-- Chelidae Gray, 1825 [Chelyidae] (käärmekaulakilpikonnat) `--o Pelomedusoides Cope, 1868 sensu Broin, 1988 [Pelumedusidae sensu lato] |--o Pelomedusera Cope, 1868 sensu Gaffney, Tong & Buffetaut, 2006 [Pelomedusoidea Cope, 1868 sensu Lindholm, 1929] | |?-o †Araripemydidae Price, 1973 | | |?- †Taquatomys decorata Broin, 1980; U. Cret. NAf. [not recognized by Gaffney, Tong & Buffetaut (2006); nomen dubium sensu Sereno & El Shafie, 2012] | | |-- †Laganemys tenerensis Sereno & El Shafie, 2012 | | `--o †Araripemys Price, 1973; U.Cret. NSA. NAf. | | |-- †A. barretoi Price, 1973 | | |-- †A. sp.2 [North Africa] | | `-- †A. sp.3 [North Africa] | `-- Pelomelusidae Cope, 1868 [Eusarkiidae] (piilokaulakilpikonnat) `--o Podocnemidera Cope, 1868 |?- †Teneremys lapparenti Broin, 1980 |--o †Euraxemydoidea Gaffney, Tong & Buffetaut, 2006 | `--o †Euraxemydidae Gaffney, Tong & Buffetaut, 2006 | |-- †Euraxemys essweini Gaffney, Tong & Buffetaut, 2006 | `-- †Dirqadim schaefferi Gaffney, Tong & Buffetaut, 2006 `--o Podocnemidoidea Cope, 1868 sensu Gaffney, Tong & Buffetaut, 2006 [Podocnemoidea Cope, 1868 sensu Meylan, 1996] |--o †Bothremydinura Baur, 1891 sensu Gaffney, Tong & Buffetaut, 2006 | `-- †Bothremydidae Baur, 1891 `--o Podocnemidinura Cope, 1868 sensu Gaffney, Tong & Buffetaut, 2006 [Podocnemoidae] |?- †Cambaremys langertoni de França & Langer, 2005; L.Cret. SAm. |-- †Brasilemys josai Lapparent de Broin, 2000a `--+-- †Portezueloemys patagonica `--+-- †Hamadachelys escuilliei Tong & Buffetaut, 1996 |--o †Peiropemydidae Gaffney, Meylan, Wood, Simons and Campos, 2011 sensu Ferreira, Iori, Hermanson & Langer, 2018 [nomen translatum ex Peiropemydodda Gaffney, Meylan, Wood, Simons and Campos, 2011] | |-- †Yuraramirim montealtensis Ferreira, Iori, Hermanson & Langer, 2018 | `--o Peiropemydodda Gaffney, Meylan, Wood, Simons and Campos, 2011 | |-- †Peiropemys mezzaliraiGaffney, Meylan, Wood, Simons and Campos, 2011 | `--+-- †Lapparentemys vilavilensis (de Broin, 1971) | `-- †Pricemys caiera `-- Podocnemidae Cope, 1868 [Podocnemididae sensu Gaffney et al., 2011] (arraukilpikonnat)
Pleurodira incertae sedis:
†Apoidochelys
†Palaeaspis [Palaeochelys, Palemys]
†Potamochelys
After Ferreira, Bronzati, Langer & Sterli, 2018<==o Pan-Pleurodira |--+-- †Platychelys | `-- †Notoemys `--o Pleurodira |-- Chelidae `--+--+--o †Araripemydidae | | |-- †Araripemys | | `-- †Laganemys | `--o †Euraxemydidae | |-- †Euraxemys | `-- †Dirqadim `--o Pelomedusoides |-- Pelomedusidae `--+-- †Sokatra `--+-- †Atolchelys `--+-- †Bothremydidae `--+-- †Brasilemys `--+-- †Hamadachelys `--+-- †Portezueloemys `--+-- †Cambaremys `--+-- †Bauruemys `--+--o †Peiropemydidae | |-- †Peiropemys | `--+-- †lapparentemys | `-- †Pricemys `-- Podocnemididae Reference(s):
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After Cadena, Bloch & Jaramillo, 2010<==o Panpelomedusoides |-- †Araripemys barretoi `--+-- †Euraxemys essweini `--o Pelomedusoides |-- Pelomedusidae `--o Panpodocnemididae |-- †Bothremydidae `--o Podocnemoidae |-- †Brasilemys josai `--+-- †Hamadachelys escuilliei |-- †Portezueloemys patagonica `--o Podocnemididae |-- †Bauruemys elegans |-- aff. †Roxochelys vilavilensis |--+-- †Cerrejonemys wayuunaiki | `-- Podocnemis `--+-- †Neochemys arenarum |-- †Neochemys laurenti [= †Papoulemys laurenti] |--+-- Erymnochelys madagascariensis | `-- Peltocephalus dumerilianus `--+-- †Decquemys paleomorpha `--+-- †Shweboemys pilgrimi |-- †Stereogenys cromeri `-- †Bairdemys Reference(s):
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Reference(s):
- Ferreira, G. S., Bronzati, M., Langer, M. C. & Sterli, J., 2018: Phylogeny, biogeography and diversification patterns of side-necked turtles (Testudines: Pleurodira).
–Royal Society Open Science: Vol. 5, #3, Art.nr. 171773, pp. (1-17) [doi: 10.1098/rsos.171773] - Ferreira, G. S., Iori, F. V., Hermanson, G. & Langer, M. C., 2018: New turtle remains from the Late Cretaceous of Monte Alto-SP, Brazil, including cranial osteology, neuroanatomy and phylogenetic position of a new taxon.
–PalZ: in press [doi: 10.1007/s12542-017-0397-x] - de la Fuente, M. S. & Iturralde-Vinent, M., 2001: A new pleurodiran turtle from the Jagua Formation (Oxfordian) of Western Cuba.
–Journal of Paleontology: Vol. 75, #4, pp. 860-869 - Gaffney, E. S., Meylan, P. A., Wood, R. C., Simons, E. & De Almeida Campos, D., 2011: Evolution of the Side-Necked Turtles: The Family Podocnemididae.
–Bulletin of the American Museum of Natural History: Vol. #350, Art.nr. pp. (1-237) [doi: 10.1206/350.1] - Gaffney, E. S., Tong, H. & Buffetaut, E., 2006: Evolution of the side-necked turtles: the families Bothremydidae, Euraxemydidae, and Araripemydidae.
–Bulletin of the American Museum of Natural History: #300, pp. 1-698 - Lapparent de Broin, F. de, 2000: African chelonians from the Jurassic to the Present: Phases of development and preliminary catalogue of the fossil record.
–Palaeontologica Africana:Vol. 36, pp. 43-82 - Meylan, P. A., 1996: Skeletal morphology and relationships of the Early Cretaceous side-necked turtle, Araripemys barretoi (Testudines: Pelomedusoides: Araripemydidae), from the Santana Formation of Brazil.
–Journal of Vertebrate Paleontology: Vol. 16, #1, pp. 20-23 - Maylan, P. A., 2001: Testudines – Turtles, tortoises and terrapins
–iNet: The Tree of Life Project: http://www.tolweb.org/tree?group=Testudines&contgroup=Amniota - Meylan, P. A. & Ganko, E., 1997: Higher Relationships of Testunides – The extinct and living turtles.
–iNet: The Tree of Life Project: [not available] - Meylan, P. A. & Ganko, E., 1997: Pleurodira – The Side-necked turtles.
–iNet: The Tree of Life Project: [not available] - Pérez-García, A., Gasulla, J. M. & Ortega, F., 2014: Eodortoka morellana gen. et sp. nov., the first pan-pleurodiran turtle (Dortokidae) defined in the Lower Cretaceous of Europe.
–Cretaceous Research: Vol. 48, pp. 130-138 [doi: http://dx.doi.org/10.1016/j.cretres.2013.12.004] - Uetz, P., Etzold, T. & Chenna, R., (eds.), 1995-2000: The Reptile Database [former “EMBL Reptile Database”].
–iNet: http://www.reptile-database.org/