After Hurum & Sabath, 2003; Currie, 2003, Currie, Hurum & Sabath, 2003, Carr, Williamson & Schwimmer, 2005, Xu, Clark, Forster, Norell, Erickson, Eberth, Jia & Zhao, 2006, Rauhut, Milner & Moore-Fay, 2009, Sereno, Tan, Brusatte, Kriegstein, Zhao & Cloward, 2009, and Brusatte, Norell, Carr, Erickson, Hutchinson, Balanoff, Bever, Choiniere, Makovicky & Xu, 2010

Alternative phylogeny after Porfiri, Novas, Calvo, Agnolín, Ezcurra & Cerda, 2014

Alternative phylogeny after Fiorillo & Tykoski, 2014

Alternative phylogeny after Loewen, Irmis, Sertich, Currie & Sampson, 2013

Alternative phylogeny after Brusatte & Benson, 2013

Alternative phylogeny after Sereno, Tan, Brusatte, Kriegstein, Zhao & Cloward, 2009, and Brusatte, Norell, Carr, Erickson, Hutchinson, Balanoff, Bever, Choiniere, Makovicky & Xu, 2010

Alternative phylogeny after Holtz, 2000, and Carr, Williamson & Schwimmer, 2005

<==o †Tyrannosauroidea Osborn, 1906 sensu Currie, Hurum & Sabath, 2003 [= Deinodontoidea]
   |--o †Proceratosauridae Rauhut, Milner & Moore-Fay, 2010 sensu Averianov et al., 2010 [Proceratosauridae Averianov, Krasnolutskii & Ivantsov, 2010]
   |  |-- †Kileskus aristotocus Averianov, Krasnolutskii & Ivantsov, 2010; M.Jur. NCAs.
   |  `--+-- †Guanlong wucaii Xu, Clark, Forster, Norell, Erickson, Eberth, Jia & Zhao, 2006; L.Jur. EAs.
   |     `--+-- †Proceratosaurus bradleyi (Woodward, 1910) Huene, 1926a [Megalosaurus bradleyi Woodward, 1910, Proceratosaurus bradleyi Huene, 1926a]; M.Jur. WEu.
   |        `-- †Sinotyrannus kazuoensis Ji Q., Ji S.-a., Zhang, L.-j., 2009; E.Cret. EAs.
   `--+-- †Dilong paradoxus Xu, Norell, Kuang, Wang, Zhao & Jia, 2004; L.Cret. EAs.
      `--+-- †Yutyrannus huali Xu, K. Wang, Zhang, Ma, Xing, Sullivan, Hu, Cheng & S. Wang, 2012; E.Cret. EAs.
         `--+-- †Timurlengia euotica Brusatte, Averianov, Sues, Muir & Butler, 2016; L.Cret. EAs.
            |--+-- †Stokesosaurus clevelandi Madsen, 1974; L.Jur. WNAm.
            |  `--+-- †Eotyrannus lengi Hutt, Naish, Martill, Barker & Newbery, 2001; L.Cret. WEu.
            |     `-- †Juratyrant langhami (Benson, 2008) Brusatte & Benson, 2013 [Stokesosaurus langhami Benson, 2008] L.Jur. WEu.
            `--+-- †Xiongguanlong baimoensis Li, Norell, Gao, Smith & Makovicky, 2009; E.Cret. CAs.
               `--+-- †Dryptosaurus aquilungis (Cope, 1866) [Lealaps aquilungis Cope, 1866]; L.Cret. ENAm.
                  `--+?- †Raptorex kriegsteini Sereno, Tan, Brusatte, Kriegstein, Zhao & Cloward, 2009; E.Cret. EAs. [juveline]
                     `--+-- †Appalachiosaurus montgomeriensis Carr, Williamson & Schwimmer, 2005; L.Cret. ENAm.
                        `--+-- †Bistahieversor sealeyi Carr & Williamson, 2010; L.Cret. SWNAm.
                           `--o †Tyrannosauridae Osborn, 1906 [=? Deinodontidae Leidy, 1856]
                              |--o †Albertosaurinae
                              |  |-- †Gorgosaurus libratus Lambe, 1914c; L.Cret. WNAm. [taxonomic history]
                              |  `-- †Albertosaurus sarcophagus Osborn, 1905 sensu Russell, 1970; L.Cret. WNAm. [taxonomic history]
                              `--o † †Tyrannosaurinae (Osborn, 1906) Matthew & Brown, 1922 [= Deinodontinae]
                                 |--o †Alioramini Olskevsky, 1995 sensu Lü, Yi, Brusatte, Yang, Li & Chen, 2014
                                 |  |-- †Qianzhousaurus sinensis Lü, Yi, Brusatte, Yang, Li & Chen, 2014; L.Cret. EAs.
                                 |  `--o †Alioramus Kurzanov, 1976b
                                 |     |-- †A. remotus Kurzanov, 1976b; L.Cret. EAs.
                                 |     `-- †A. altai Brusatte, Carr, Erickson, Bever & Norell, 2009; L.Cret. EAs.
                                 `--+-- †Teratophoneus curriei Carr et al., 2011
                                    `--+-- †Lythronax argestes Loewen, Irmis, Sertich, Currie & Sampson, 2013 L.Cret. WNAm.
                                       `--+?- †Zhuchengtyrannus magnus Hone, Wang, Sullivan, Zhao, Chen, Li, Ji, Ji & Xu, 2011; L.Cret. EAs.
                                          |?- †Nanuqsaurus hoglundi Fiorillo & Tykoski, 2014; L.Cret. NWNAm.
                                          |--o †Tarbosaurini Olshevsky & Ford, 1995
                                          |  |-- †Tarbosaurus bataar (Maleev, 1955a) Maleev, 1955b [taxonomic history] L.Cret. EAs. (“mongoliantyrannosaurus”)
                                          |  `--o †Daspletosaurus Russell, 1970a
                                          |     |-- †D. sp. indet. [L.Cret. WNAm.] [Currie, 2003]
                                          |     |-- †D. sp. indet. [L.Cret. WNAm.] [Currie, 2003]
                                          |     `-- †D. torosus Russell, 1970a; L.Cret. NAm. (“härkätyrannisaurus”)
                                          `--o †Tyrannosaurini Osborn, 1906
                                             |?- †Nanotyrannus lancensis (Gilmore, 1946c) Bakker, et al., 1988 L.Cret. WNAm. [probably juveline T. rex]
                                             `-- †Tyrannosaurus rex Osborn, 1905 [taxonomic history] L.Cret. WNAm. (kuningastyrannisaurus)

Other possible Tyrannosauroids:

Albertosaurus periculosus Riabinin, 1930 (nomen dubium, possibly Tarbosaurus)
    †Alectrosaurus olseni Gilmore, 1933a (nomen dubium)
    †Aviatyrannis jurassica Rauhut, 2003; U.Jur. SWEu.
    †Bagaraatan ostromi Osmolska, 1996
    †Chingkankousaurus fragilis Young, 1958
    †Deinodon Leidy, 1856 (nomen vanum sensu Russell, 1970)
    †Dinotyrannus megagracilis (Paul, 1988) [Albertosaurus megagracilis Paul, 1988]
    †Dryptosaurus aquilungis (Cope, 1866) [Lealaps aquilungis Cope, 1866]
    †Labocania anomola Molnar, 1974
    †Nuthetes destructor Owen, 1854 [Megalosaurus destructor (Owen, 1854) Steel, 1970]
    †Prodeinodon mongoliensis Osborn, 1924 (incertae sedis)
    †Prodeinodon kwangshiensis Hou, et al., 1975
    ?†Santanaraptor placidus Kellner, 1999; E.Cret. NESAm. [also treated as velocisaurid abelisauroid]
    ?†Tanycolagreus topwilsoni Carpenter, Miles & Cloward, in Carpenter, 2005 [also treated as coelurid basal coelurosaurian]
    †Timimus hermani Rich & Vickers-Rich, 1994; E.Cret. SEAust. [sensu Benson, Rich, Vickers-Rich & Hall, 2012]
    †Tyrannosaurus luanchuanensis Dong, 1979 (“kiinantyrannisaurus”)
    NMV-P186069 [isolated pubis from E.Cret. Victoria, Australia] [Benson et al., 2010]

    †Megaraptora Benson, Carrano & Brusatte, 2009 [has been treated as Neovenatorid allosauroids]
        ?†Siats meekerorum Zanno & Makovicky, 2013; L.Cret. NAm.
        †Fukuiraptor kitadaniensis Azuma & Currie, 2000
        †Megaraptoridae Novas, Agnolín, Ezcurra, Porfiri & Canale, 2013
            †Australovenator wintonensis Hocknull et al., 2009
            †Aerosteon riocoloradensis Sereno et al., 2009
            †Megaraptor namunhuaiquii Novas, 1998
            †Orkoraptor burkei Novas, Ezcurra & Lecuona, 2008

Alternative phylogenies of Tyrannosauroidea:

After Porfiri, Novas, Calvo, Agnolín, Ezcurra & Cerda, 2014

<==o †Tyrannosauroidea
   |--o †Proceratosauridae
   |  |-- †Dilong
   |  `--+-- †Tanycolagreus
   |     `--+-- †Guanlong
   |        |-- †Kileskus
   |        `-- †Proceratosaurus
   `--+-- †Santanaraptor
      `--+--o †Megaraptora
         |  |-- †Fukuiraptor
         |  `--o †Megaraptoridae
         |     |-- †Australovenator
         |     |-- †Aerosteon
         |     |-- †Megaraptor
         |     `-- †Orkoraptor
         `--+-- †Xiongguanlong
            `--+-- †Appalachiosaurus
               `--o †Tyrannosauridae
                  |-- †Albertosaurus
                  `-- †Tyrannosaurus

Reference(s):

  • Porfiri, J. D., Novas, F. E., Calvo, J. O., Agnolín, F. L., Ezcurra, M. D. & Cerda, I. A., 2014: Juvenile specimen of Megaraptor (Dinosauria, Theropoda) sheds light about tyrannosauroid radiation.
    –Cretaceous Research: Vol. 51, pp. 35-55 [doi: http://dx.doi.org/10.1016/j.cretres.2014.04.007]

Fiorillo & Tykoski, 2014

This phylogeny is based on Loewen et al. datamatrix with Nanuqsaurus added.

<==o †Tyrannosauridae
   |--+-- †Albertosaurus
   |  `-- †Gorgosaurus
   `--o †Tyrannosaurinae
      |--+-- †Daspletosaurus
      |  `-- “Two Medicine OTU”
      `--+-- †Teratophoneus
         `--+-- †Bistahieversor
            `--+-- †Lythronax
               `--+-- †Nanuqsaurus
                  `--+-- †Tyrannosaurus
                     `--+-- †Tarbosaurus
                        `-- †Zhuchengtyrannus

Reference(s):

  • Fiorillo, A. R. & Tykoski, R. S., 2014: A Diminutive New Tyrannosaur from the Top of the World.
    –Plos One: Vol. 9, #3, pp. e91287 (1-13) [doi: 10.1371/journal.pone.0091287]

After Loewen, Irmis, Sertich, Currie & Sampson, 2013

<==o †Tyrannosauroidea
   |--o †Proceratosauridae
   |  |--+-- †Proceratosaurus
   |  |  |-- †Kileskus
   |  |  `-- †Guanlong
   |  `--+-- †Sinotyrannus
   |     `--+-- †Juratyrant
   |        `-- †Stokesosaurus
   `--+-- †Dilong
      `--+-- †Eotyrannus
         `--+-- †Bagaraatan
            `--+-- †Raptorex
               `--+-- †Dryptosaurus
                  `--+--+-- †Alectrosaurus
                     |  `-- †Xiongguanlong
                     `--+-- †Appalachiosaurus
                        `--+-- †Alioramus
                           `--o †Tyrannosauridae
                              |--+-- †Albertosaurus
                              |  `-- †Gorgosaurus
                              `--o †Tyrannosaurinae
                                 |-- “Dinosaur Park OTU”
                                 `--+-- †Daspletosaurus
                                    `--+-- “Two Medicine OTU”
                                       `--+-- †Teratophoneus
                                          `--+-- †Bistahieversor
                                             `--+-- †Lythronax
                                                `--+-- †Tyrannosaurus
                                                   `--+-- †Tarbosaurus
                                                      `-- †Zhuchengtyrannus

Reference(s):

  • Loewen, M. A., Irmis, R. B., Sertich, J. J. W., Currie, P. J. & Sampson, S. D., 2013: Tyrant Dinosaur Evolution Tracks the Rise and Fall of Late Cretaceous Oceans.
    –Plos One: Vol. 8, #11, pp. (e79420) [doi: 10.1371/journal.pone.0079420]

After Brusatte & Benson, 2013

<==o †Tyrannosauroidea
   |--o †Proceratosauridae
   |  |-- †Kileskus
   |  `--+-- †Guanlong
   |     `-+-- †Proceratosaurus
   |       `-- †Sinotyrannus
   `--+-- †Dilong
      `--+--+-- †Stokesosaurus
         |  `--+-- †Eotyrannus
         |     `-- †Juratyrant
         `--+-- †Xiongguanlong
            `--+-- †Dryptosaurus
               `--+-- †Raptorex
                  `--+-- †Appalachiosaurus
                     `--+-- †Bistahieversor
                        `--o †Tyrannosauridae
                           |--+-- †Albertosaurus
                           |  `-- †Gorgosaurus
                           `--+-- †Alioramus
                              `--+-- †Teratophoneus
                                 `--+-- †Daspletosaurus
                                    `--+-- †Tarbosaurus
                                       `-- †Tyrannosaurus

Reference(s):

  • Brusatte, S. L. & Benson, R. B. J., 2013: The systematics of Late Jurassic tyrannosauroid theropods from Europe and North America.
    –Acta Palaeontologica Polonica: Vol. 58, #1, pp. 47-54 [doi: 10.4202/app.2011.0141]

After Sereno, Tan, Brusatte, Kriegstein, Zhao & Cloward, 2009, and Brusatte, Norell, Carr, Erickson, Hutchinson, Balanoff, Bever, Choiniere, Makovicky & Xu, 2010

<==o †Tyrannosauroidea
   |--o †Proceratosauridae
   |  |-- †Kileskus
   |  `--+-- †Guanlong
   |     `-+-- †Proceratosaurus
   |       `-- †Sinotyrannus
   `--+-- †Dilong
      `--+--+-- †Eotyrannus
         |  `-- †Stokesosaurus
         `--+-- †Xiongguanlong
            `--+-- †Dryptosaurus
               `--+-- †Raptorex
                  `--+-- †Appalachiosaurus
                     `--+-- †Bistahieversor
                        `--o †Tyrannosauridae
                           |--+-- †Albertosaurus
                           |  `-- †Gorgosaurus
                           `--+-- †Alioramus
                              `--+-- “Utah taxon”
                                 `--+-- †Daspletosaurus
                                    `--+-- †Tarbosaurus
                                       `-- †Tyrannosaurus

Reference(s):

  • Brusatte, S. L., Carr, T. D., Erickson, G. M., Bever, G. S. & Norell, M. A., 2009: A long-snouted, multihorned tyrannosaurid from the Late Cretaceous of Mongolia.
    –Proceedings of the National Academy of Sciences: Vol. 106, #41, pp. 17261-17266 [doi: 10.1073/pnas.0906911106]
  • Brusatte, S. L., Norell, M. A., Carr, T. D., Erickson, G. M., Hutchinson, J. R., Balanoff, A. M., Bever, C. S., Choiniere, J. N., Makovicky, P. J. & Xu, X., 2010: Tyrannosaur Paleobiology: New Research on Ancient Exemplar Organisms.
    –Science: Vol. 329, pp. 1481-1485 [doi: 10.1126/science.1193304]
  • Sereno, P. C., Tan, L., Brusatte, S. L., Kriegstein, H. J., Zhao, X. & Cloward, K., 2009: Tyrannosaurid skeletal design first evolved at small body size.
    –Science: Vol. 326, pp. 418-422 (+ supplement 1-27) [doi: 10.1126/science.1177428]

After Holtz, 2000, and Carr, Williamson & Schwimmer, 2005 (reduced to genera)

<==o †Tyrannosauroidea
   |-- †Guanlong
   `--+-- †Dilong
      `--+-- †Eotyrannus
         `--+?- †Siamotyrannus
            |?- †Labocania
            |--+?- †Stokesosaurus
            |  `?- †Itemirus
            `--+?- †Dryptosaurus
               `--+?- †Appalachiosaurus
                  `--o †Tyrannosauridae
                     |?- †Bistahieversor
                     |==o †“Aublysodontinae” [juvelines]
                     |  |?- †“Kirtland Shale aublysodontine”
                     |  |?- †Aublysodon
                     |  |?- †Alectrosaurus
                     |  `?- †Stygivenator [Aublysodon]
                     `--o †Tyrannosaurinae
                        |?- †“Two Medicine tyrannosaurine”
                        |-- †Alioramus
                        `--+?- †Shanshanosaurus
                           `--+-- †Albertosaurus
                              |-- †Gorgosaurus
                              `--o ?†Tyrannosaurini
                                 |-- †Daspletosaurus
                                 `--+-- †Tarbosaurus
                                    `-- †Tyrannosaurus

Reference(s):

  • Carr, T. D., Williamson, T. E. & Schwimmer, D. R., 2005: A new genus and species of tyrannosauroid from the Late Cretaceous (Middle Campanian) Demopolis Formation of Alabama.
    –Journal of Vertebrate Paleontology: Vol. 25, #1, pp. 119-143
  • Holtz, T. R. Jr., 2000: †Tyrannosauroidea – Tyrant dinosaurs and their possible relatives.
    –iNet: The Tree of Life
  • Holtz, T. R. Jr., 2000: †Tyrannosauridae – Tyrant dinosaurs.
    –iNet: The Tree of Life

Reference(s):