After Holtz, 1998, 1999, 2001, Xu, Norell, Wang, Makovicky & Wu, 2002, Clark, Norell & Makovicky, 2002, Hwang, Norell, Ji & Gao, 2002, Novas & Pol, 2002, Naish, Martill & Frey, 2004, and Rauhut & Xu, 2005
Alternative, older phylogeny of coelurosaurs.
<==o Coelurosauria von Huene, 1914 sensu Holtz, 1999 |?- †Aorun zhaoi Choiniere, Clark, Forster, Norell, Eberth, Erickson, Chu & Xu, 2014 |?- †Bahariasaurus ingens Stromer, 1934a |?- †Lourinhanosaurus antunesi Mateus, 1998 |?- †Ozraptor subotaii Long & Molnar, 1998 [an abelisauroid?] |?- †Gasosaurus constructus Dong & Tang, 1985 [?Kaijiangosaurus lini He, 1984] |?- †Zuolong salleei Choiniere, Clark, Forster & Xu, 2010 `--+-- †Tugulusaurus faciles Dong, 1973 [taxonomic history] |--o †Ornitholestidae Paul, 1988a [Allosauridae: Ornitholestinae Paul, 1988a] | |?- †Rapator ornitholestoides Huene, 1932 | |?- †Piveteausaurus divesensis (Walker, 1964) [Eustreptospondylus divesensis Walker, 1964 partim] [see Spinosauroidea] | |?- †Proceratosaurus bradleyi (Woodward, 1910) sensu Huene, 1926a [Megalosaurus bradleyi Woodward, 1910, Proceratosaurus bradleyi Huene, 1926a] [a basal tyrannosauroid sensu Rauhut, Milner & Moore-Fay, 2009] | |?- †Nedcolbertia justinhoffmanni Kirkland, Britt, Whittle, Madsen & Burge, 1998 ["Nedcolbertia" whittlei] [a basal ornithomimosaur aensu Brownstein, 2017] | `-- †Ornitholestes hermanni Osborn, 1903 ("linnunnappaaja") `--+--o †Coeluridae Marsh, 1881 | |-- †Coelurus fragilis March, 1879c [Coelurus agilis Marsh, 1884a] | `-- †Tanycolagreus topwilsoni Carpenter, Miles & Cloward, in Carpenter, 2005 `--+-- †Scipionyx samnicus Del Sasso, & Signore, 1998 `--+?- †Aniksosaurus darwini Martínez & Novas, 2006 |?-o †Compsognathidae Marsh, 1882 (compsognathids; siroleuat) | |?- †Sinocalliopteryx gigas S. Ji, Q. Ji, Lü & Yuan, 2007 | |-- †Juravenator starki Göhlich & Chiappe, 2006 [“Compsognathus” sp.] U.Jur. CEu. | |-- †Huaxiagnathus orientalis Hwang, Norell, Ji & Gao, 2004; L.Cret. EAs. | `--+-- †Sinosauropteryx prima Ji & Ji, 1996; L.Cret. EAs. [feathered] | `--+-- †Mirischia asymmetrica Naish, Martill & Frey, 2004; L.Cret. ESA. | `--+?-o †Aristosuchus Seeley, 1887 [nomen dubium] | | |-- †A. pusillus (Fox, 1876) | | `?- †A. oweni (Fox, 1866) | `-- †Compsognathus longipes Wagner, 1861 [incl. Compsognathus corallestris Bidar, Demay & Thomel, 1972] L.Cret. EAs. `--+-- †Tyrannosauroidea Osborn, 1906 sensu Holtz, 1998 `--o Maniraptoriformes [Proavia Paul, 1988] |?- †Archaeornithoides deinosauriscus Elzanowski & Wellnhofer, 1992 |?- †Bradycneme draculae Harrison & Walker, 1975 |?- †Kakuru kujani Molnar & Pledge, 1980 |?- †Koparion douglassi Chure, 1994 `--+-- †Ornithomimosauria Barsbold, 1976b `--o Maniraptora Gauthier, 1986 |?- †Fukuivenator paradoxus Azuma, Xu, Shibata, Kawabe, Miyata & Imai, 2016; E.Cret. EAs. |?- †Yixianosaurus longimanus Xu & Wang, 2003; E.Cret. EAs. |?-o †Richardoestesia Currie, Rigby & Sloan, 1990 | |-- †R. gilmorei Currie, Rigby & Sloan, 1990 | `-- †R. isosceles Sankey, 2001 `--o Avepectora Paul, 2002 `--o Metornithes sensu Xu, Norell, Wang, Makovicky & Wu, 2002 |--o †Alvarezsauria Livezay & Zusi, 2007 [Alvarezsauroidea Choiniere, Xu, Clark, Forster, Guo & Han, 2010] | `-- †Alvarezsauridae Bonaparte, 1991 sensu Xu, Norell, Wang, Makovicky & Wu, 2002, and Novas & Pol, 2002 `--+--+-- Paraves sensu Xu, Norell, Wang, Makovicky & Wu, 2002 [Avialae, Troodontidae & Dromaeosauridae] (linnut ja sirppikynnet; birds & near-birds) | `--+?- †Bagaraatan ostromi Osmólska, 1996 | `-- †Xinjiangovenator parvus Rauhut & Xu, 2005 [taxonomic history] `--o †Oviraptoriformes Sereno |?- †Microvenator celer Ostrom, 1970a |?- †Therizinosauroidea Maleev, 1954 sensu Russell & Dong, 1993 [Segnosauria Perle, 1979 sensu Barsbold & Perle, 1980; Segnosaurischia] (segnosaurit; segnosaurs, therizinosaurs) `-- †Oviraptorosauria Barsbold, 1976b (oviraptors; oviraptorit)
Coelurosauria incertae sedis:
†Phaedrolosaurus ilikensis Dong, 1973 (nomen dubium) [taxonomic history]
After Xu, Zheng, Sullivan, Wang, Xing, Wang, Zhang, O'Connor, Zhang & Pan, 2015<==o Coelurosauria |--+-- †Tyrannosauroidea | `--o †Coeluridae | |-- †Coelurus | `-- †Tanycolagreus `--+--o †Compsognathidae | |-- †Huaxiagnathus | `--+-- †Sinosauropteryx | `-- †Compsognathus `--+-- †Ornithomimosauria `--+-- †Ornitholestes `--+-- †Alvarezsauridae `--+-- †Therizinosauroidea `--+-- †Protarchaeopteryx `--+-- †Oviraptorosauria `-- Paraves Reference(s):
|
After Porfiri, Novas, Calvo, Agnolín, Ezcurra & Cerda, 2014<==o Coelurosauria |-- †Chilantaisaurus `--+--o †Compsognathidae | |-- †Compsognathus | `-- †Sinosauropteryx |--+-- †Dromaeosauridae | |-- †Oviraptorosauria | `-- †Ornithomimosauria `--o †Tyrannosauroidea |--o †Proceratosauridae | |-- †Dilong | `--+-- †Tanycolagreus | `--+-- †Guanlong | |-- †Kileskus | `-- †Proceratosaurus `--+-- †Santanaraptor `--+--o †Megaraptora | |-- †Fukuiraptor | `--o †Megaraptoridae | |-- †Australovenator | |-- †Aerosteon | |-- †Megaraptor | `-- †Orkoraptor `--+-- †Xiongguanlong `--+-- †Appalachiosaurus `--o †Tyrannosauridae |-- †Albertosaurus `-- †Tyrannosaurus Reference(s):
|
Alternative, older phylogeny of coelurosaurs<==o Coelurosauria `--o Maniraptoriformes |?-o †Coeluridae | |-- †Coelurus | `-- †Tanycolagreus |?-o †Compsognathidae | |-- †Juravenator | |-- †Huaxiagnathus | `--+-- †Sinosauropteryx | `--+-- †Mirischia | `--+?- †Aristosuchus | `-- †Compsognathus `--+--o †Arctometatarsalia | |-- †Tyrannosauroidea | `--o †Bullatosauria | |-- †Troodontidae | `-- †Ornithomimosauria `--o Maniraptora |-- †Ornitholestidae `--+--o Eumaniraptora | |-- †Dromaeosauridae | `--o Avialae | `-- Aves `--o †Oviraptoriformes |?- †Microvenator |?- †Therizinosauroidea `-- †Oviraptorosauria |
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