After Holtz, 1998, 1999, 2001, Xu, Norell, Wang, Makovicky & Wu, 2002, Clark, Norell & Makovicky, 2002, Hwang, Norell, Ji & Gao, 2002, Novas & Pol, 2002, Naish, Martill & Frey, 2004, and Rauhut & Xu, 2005
Alternative, older phylogeny of coelurosaurs.
<==o Coelurosauria von Huene, 1914 sensu Holtz, 1999 |?- †Bahariasaurus ingens Stromer, 1934a |?- †Nqwebasaurus thwazi de Klerk et al., 2000 |?- †Ozraptor subotaii Long & Molnar, 1998 [an abelisauroid?] |?- †Gasosaurus constructus Dong & Tang, 1985 [?Kaijiangosaurus lini He, 1984] `--+-- †Tugulusaurus faciles Dong, 1973 [taxonomic history] |-- †Ornitholestidae Paul, 1988a `--+--o †Coeluridae Marsh, 1881 | |-- †Coelurus fragilis March, 1879c [Coelurus agilis Marsh, 1884a] | `-- †Tanycolagreus topwilsoni Carpenter, Miles & Cloward, in Carpenter, 2005 `--+-- †Scipionyx samnicus Del Sasso, & Signore, 1998 `--+?-o †Compsognathidae Marsh, 1882 (compsognathids; siroleuat) | |?- †Sinocalliopteryx gigas S. Ji, Q. Ji, Lü & Yuan, 2007 | |-- †Juravenator starki Göhlich & Chiappe, 2006 [“Compsognathus” sp.] U.Jur. CEu. | |-- †Huaxiagnathus orientalis Hwang, Norell, Ji & Gao, 2004; L.Cret. EAs. | `--+-- †Sinosauropteryx prima Ji & Ji, 1996; L.Cret. EAs. [feathered] | `--+-- †Mirischia asymmetrica Naish, Martill & Frey, 2004; L.Cret. ESA. | `--+?-o †Aristosuchus Seeley, 1887 [nomen dubium] | | |-- †A. pusillus (Fox, 1876) | | `?- †A. oweni (Fox, 1866) | `-- †Compsognathus longipes Wagner, 1861 [incl. Compsognathus corallestris Bidar, Demay & Thomel, 1972] L.Cret. EAs. `--+-- †Tyrannosauroidea Osborn, 1906 sensu Holtz, 1998 `--o Maniraptoriformes [Proavia Paul, 1988] |?- †Archaeornithoides deinosauriscus Elzanowski & Wellnhofer, 1992 |?- †Bradycneme draculae Harrison & Walker, 1975 |?- †Kakuru kujani Molnar & Pledge, 1980 |?- †Koparion douglassi Chure, 1994 `--+-- †Ornithomimosauria Barsbold, 1976b `--o Maniraptora Gauthier, 1986 |?-o †Richardoestesia Currie, Rigby & Sloan, 1990 | |-- †R. gilmorei Currie, Rigby & Sloan, 1990 | `-- †R. isosceles Sankey, 2001 `--o Avepectora Paul, 2002 `--o Metornithes sensu Xu, Norell, Wang, Makovicky & Wu, 2002 |-- †Alvarezsauridae sensu Xu, Norell, Wang, Makovicky & Wu, 2002 and Novas & Pol, 2002 `--+--+-- Paraves sensu Xu, Norell, Wang, Makovicky & Wu, 2002 [Avialae, Troodontidae & Dromaeosauridae] (linnut ja sirppikynnet; birds & near-birds) | `--+?- †Bagaraatan ostromi Osmólska, 1996 | `-- †Xinjiangovenator parvus Rauhut & Xu, 2005 [taxonomic history] `--o †Oviraptoriformes Sereno |?- †Microvenator celer Ostrom, 1970a |?- †Therizinosauroidea Maleev, 1954 sensu Russell & Dong, 1993 [Segnosauria Perle, 1979 sensu Barsbold & Perle, 1980; Segnosaurischia] (segnosaurit; segnosaurs, therizinosaurs) `-- †Oviraptorosauria Barsbold, 1976b (oviraptors; oviraptorit)
Coelurosauria incertae sedis:
†Phaedrolosaurus ilikensis Dong, 1973 (nomen dubium) [taxonomic history]
Reference(s):
- Carpenter, K., Miles, C. A. ∓ Cloward, K., 2005: 2. New small theropod from the Upper Jurassic Morrison Formation of Wyoming. 23-48.
in Carpenter, K., (eds.) 2005: The Carnivorous Dinosaurs.
–Indiana University Press, Bloomington, 2005 - Chen, P.-J., Dong, Z.-M. & Zhen, S.-N., 1998: An exceptionally well-preserved theropod dinosaur from the Yixian Formation of China.
–Nature: Vol. 391, 8 January, pp. 147-152 - Currie, P. J. & Chen, P.-j., 2001: Anatomy of Sinosauropteryx prima from Liaoning, northeastern China.
–Canadian Journal of Earth Sciencies: Vol. 38, #12, pp. 1705-1727 - Clark, J. M., Norell, M. A. & Makovicky, P. J., 2002: Cladistic Approaches to the Relationships of Birds to Other Theropod Dinosaurs. 31-61.
in Chiappe, L. M. & Witmer, L. M., (eds.) 2002: Mesozoic Birds – Above the Heads of Dinosaurs.
–University of California Press, Berkeley, Los Angeles, London, 2002, xii-520 - Del Sasso, C. & Signore, M., 1998: Exceptional soft-tissue preservation in a theropod dinosaur from Italy.
–Nature: Vol. 392, 26 March, pp. 383-387 - Dyke, G. J. & Norell, M. A., 2005: Caudipteryx as a non-avialan theropod rather than a flightless bird.
–Acta Palaeontologica Polonica: Vol. 50, #1, pp. 101-116 - Göhlich, U. B. & Chiappe, L. M., 2006: A new carnivorous dinosaur from the Late Jurassic Solnhofen archipelago.
–Nature: Vol. 440, pp. 329-332 - Holtz, T. R., Jr., 1999: Concensus and conflict in current coelurosaur cladograms.
–Journal of Vertebrate Paleontology: Vol. 19, #3, Supplement to #3, pp. 52A - Holtz, T. R., Jr., 2001: Pedigree of the Tyrant Kings: New information on the origin and evolution of the Tyrannosauridae.
–Journal of Vertebrate Paleontology: Vol. 21, #3, Supplement to #3, pp. 62A-63A - Hwang, S. H., Norell, M. A., Ji, Q. & Gao, K.-Q., 2002: New specimens of Microraptor zhaoianus (Theropoda: Dromaeosauridae) from Northeastern China.
–American Museum Novitates: #3381, pp. 1-44 - Hwang, S. H., Norell, M. A., Ji, Q. & Gao, K.-Q., 2004: A large compsognathid from the Early Cretaceous Yixian Formation of China.
–Journal of Systematic Palaeontology: Vol. 2, #1, pp. 13-30 - Ji, S., Ji, Q., Lü, J. & Yuan, C., 2007: A New Giant Compsognathid Dinosaur with Long Filamentous Integuments from Lower Cretaceous of Northeastern China.
–Acta Geologica Sinica: Vol. 81, #1, pp. 8-15 [abstract only] - Keesey, T. M., 1996-2001: Coelurosauria - Very Advanced Theropods
–iNet: The Dinosauricon: http://dinosauricon.com/taxa/coelurosauria.html - de Klerk, W. J., Forster, C. A., Sampson, S. D., Chinsamy, A. & Ross, C. F., 2000: A new coelurosaurian dinosaur from the Early Cretaceous of South Africa.
–Journal of Vertebrate Paleontology: Vol. 20, #2, pp. 324-332 - Molnar, R., E., 1990: Problematic Theropoda: "Carnosaurs". 306-317
in Weishampel, D. B., Dodson, P., & Osmólska, H. (eds.), 1990: The Dinosauria.
–University of California Press, Berkley, Los Angeles, Oxford, 1990 xvi-733 - Naish, D., Martill, D. M. & Frey, E., 2004: Ecology, systematics and biogeographical relationships of dinosaurs, including a new theropod, from the Santana Formation (?Albian, Early Cretaceous) of Brazil.
–Historical Biology: Vol. 16, #2-4, pp. 57-70 - Norman, D., B., 1990: Problematic Theropoda: "Coelurosaurs". 280-305
in Weishampel, D. B., Dodson, P., & Osmólska, H. (eds.), 1990: The Dinosauria.
–University of California Press, Berkley, Los Angeles, Oxford, 1990 xvi-733 - Novas, F. E. & Pol, D., 2002: Alvarezsaurid Relationships Reconsidered. 121-125.
in Chiappe, L. M. & Witmer, L. M., (eds.) 2002: Mesozoic Birds - Above the Heads of Dinosaurs.
–University of California Press, Berkeley, Los Angeles, London, 2002, xii-520 - Osmólska, H., & Barsbold, R., 1990: Troodontidae. 259-268
in Weishampel, D. B., Dodson, P., & Osmólska, H. (eds.), 1990: The Dinosauria.
–University of California Press, Berkley, Los Angeles, Oxford, 1990 xvi-733 - Rauhut, O. W. M. & Xu, X., 2005: The small theropod dinosaurs Tugulusaurus and Phaedrolosaurus from the Early Cretaceous of Xinjiang, China.
–Journal of Vertebrate Paleontology: Vol. 25, #1, pp. 107-118 - Sankey, J. T., 2001: Late Campanian southern dinosaurs, Aguja Formation, Big Bend, Texas.
–Journal of Paleontology: Vol. 75, #1, pp. 208-215 - Sereno, P. C., Dutheil, D. B., Iarochene, M., Larsson, H. C. E.,Lyon, G. H., Magwene, P. M., Sidor, C. A., Verricchio, D. J. & Wilson, J. A., 1996: Predatory dinosaurs from the Sahara and Late Cretaceous faunal differentiation.
–Science: Vol. 272, 17 May, pp. 986-991 - Xu, X., Norell, M. A., Wang, X., Makovicky, P. J. & Wu, X.-C., 2002: A basal troodontid from the Early Cretaceous of China.
–Nature: Vol. 415, pp. 780-784 - Zhang, F., Zhou, Z., Xu, X. & Wang, X.-l., 2002: A juveline coelurosaurian theropod from China indicates arboreal habits.
–Naturwissenschaften: Vol. 89, #8, pp. 394-398