Mikko's Phylogeny Archive

†Therizinosauroidea – segnosaurs

After Zanno, 2010

<==o †Therizinosauria sensu Zanno, 2010 [Therizinosauroidea Maleev, 1954 sensu Russell & Dong, 1993; Segnosauria Perle, 1979 sensu Barsbold & Perle, 1980; Segnosaurischia]
   |?- †Eshanosaurus deguchiianus Xu, Zang & Clark, 2001, E. Jur. SEAs. [An Early Jurassic therizinosaur, really!?]
   |-- †Falcarius utahensis Kirkland, Zanno, Sampson, Clark & DeBlieux, 2005; E.Cret. WNAm.
   `--+-- †Jianchangosaurus yixianensis Pu, Kobayashi, Lü, Xu, Wu, Chang, Zhang & Jia, 2013; E.Cret. EAs.
      `--o †Therizinosauroidea Maleev, 1954 sensu Zanno, 2010
         |-- †Beipiaosaurus inexpectatus Xu et al., 1999, E. Cret. NEAs. [feathered]
         `--+-- †Martharaptor greenriverensis Senter, Kirkland & DeBlieux, 2012; E.Cret. NAm.
            `--+-- †Alxasaurus elesitaiensis Russell & Dong, 1995 [Alxasauridae Russell & Dong, 1995]
               `--+-- †Erliansaurus bellamanus Xu, Zhang, Sereno, Zhao, Kuang, Han & Tan, 2002
                  `--+-- †Neimongosaurus yangi Zhang, Xu, Sereno, Kwang & Tan, 2001; L.Cret. EAs.
                     `--+-- †Enigmosaurus mongoliensis Barsbold & Perle, 1983
                        `--+-- †Suzhousaurus megatheroides Daqing, Cuo, Hailu, Lamanna, Harris, Lacovara & Jianping, 2007
                           `--o †Therizinosauridae Maleev, 1954 [Segnosauridae Perle, 1979, Enigmosauridae Barsbold & Perle, 1983]
                              |-- †Erlikosaurus andrewsi Perle, 1980
                              |--o †Nanshiungosaurus Dong, 1979
                              |  |-- †N. brevispinus Dong, 1979
                              |  `?- ?†N. zheziangensis (Dong, 1979) ["Chialantaisaurus" zheziangensis Dong, 1979]
                              |--o †Nothronychus Kirkland & Wolfe vide Stanley, 2001
                              |  |-- †N. mckinleyi Kirkland & Wolfe vide Stanley, 2001
                              |  `-- †N. graffami Zanno, Gillette, Albright & Titus, 2009
                              |-- †Segnosaurus galbinensis Perle, 1979
                              `-- †Therizinosaurus cheloniformis Maleev, 1954

<==o †Therizinosauria sensu Zanno, 2010 [Therizinosauroidea Maleev, 1954 sensu Russell & Dong, 1993; Segnosauria Perle, 1979 sensu Barsbold & Perle, 1980; Segnosaurischia]
   |-- †Falcarius
   `--+-- †Beipiaosaurus
      `--+-- †Martharaptor
         `--+-- †Alxasaurus
            `--o †Therizinosauridae
               |--†Nanshiungosaurus
               `--+-- †Suzhousaurus
                  `--+-- †Nothronychus
                     `--+-- †Segnosaurus
                        `--+-- †NeimongosaurusErliansaurus
                           `--+-- †Erlikosaurus
                              `-- †Therizinosaurus

Reference(s):

• Barsbold, R., & Marya´nska, T., 1990: Segnosauria. 408-415
  in Weishampel, D. B., Dodson, P., & Osmólska, H. (eds.), 1990: The Dinosauria.
  –University of California Press, Berkley, Los Angeles, Oxford, 1990 xvi-733
• Daqing, L. I., Cuo, P., Hailu, Y. O. U., Lamanna, M. C., Harris, J. D., Lacovara, K. J. & Jianping, Z., 2007: A Large Therizinosauroid (Dinosauria: Theropoda) from the Early Cretaceous of Northwestern China.
  –Acta Geologica Sinica - English Edition: Vol. 81, #4, pp. 539-549 [doi: 10.1111/j.1755-6724.2007.tb00977.x]
• Ji, Q., Currie, P. J., Norell, M. A. & Ji, S.-A., 1998: Two feathered dinosaurs from northeastern China.
  –Nature: Vol. 393, 25 June, pp.753-761
• Keesey, T. M., 1996-2001: Therizinosauria – Weirdest of the Weird
  –iNet: The Dinosauricon: http://dinosauricon.com/taxa/therizinosauria.html
• Kirkland, J. I. & Wolfe, D. G., 2001: First definitive therizinosaurid (Dinosauria: Theropoda) from North America.
  –Journal of Vertebrate Paleontology: Vol. 21, #3, pp. 410-414
• Kirkland, J. I., Zanno, L. E., Sampson, S. D., Clark, J. M. & DeBlieux, D. D., 2005: A primitive therizinosauroid dinosaurs from the Early Cretaceous of Utah.
  –Nature: Vol. 435, pp. 84-87
• Pu, H., Kobayashi, Y., Lü, J., Xu, L., Wu, Y., Chang, H., Zhang, J. & Jia, S., 2013: An Unusual Basal Therizinosaur Dinosaur with an Ornithischian Dental Arrangement from Northeastern China.
  –Plos One: Vol. 8, #5, pp. e63423 [doi: 10.1371/journal.pone.0063423]
• Senter, P., Kirkland, J. I. & DeBlieux, D. D., 2012: Martharaptor greenriverensis, a New Theropod Dinosaur from the Lower Cretaceous of Utah.
  –Plos One: Vol. 7, #8, art.nr. e43911 [doi: 10.1371/journal.pone.0043911]
• Sereno, P. C., The evolution of dinosaurs.
  –Science: Vol. 284, 25 June, pp. 2137-2147
• Xu, X., Tang, Z.-L. & Wang, X.-L., 1999: A therizinosauroid dinosaur with integumentary structures from China.
  –Nature: Vol. 399, 27 May, pp. 350-354
• Xu, X., Zhao, X. & Clark, J. M., 2001: A new therizinosaur from the Lower Jurassic Lower Lufeng Formation of Yunnan, China.
  –Journal of Vertebrate Paleontology: Vol. 21, #3, pp. 477-483
• Xu X., Zhang X.-H., Sereno, P., Zhao, X-J., Kuang, Xu.-W., Han, J.& Tan, L., 2002: A new therizionosauroid (Dinosauria, Theropoda) from the Upper Cretaceous Iren Dabau Formation of Nei Mongol.
  –Vertebrate Palasiatica: Vol.40, #3, pp. 228-240
  [from iNet abstract at Dinodata.net (http://www.dinodata.net/Newdesuk.htm)]
• Zanno, L. E., 2010: A taxonomic and phylogenetic re-evaluation of Therizinosauria (Dinosauria: Maniraptora).
  –Journal of Systematic Palaeontology: Vol. 8, #4, pp. 503-543 [doi: 10.1080/14772019.2010.488045]
• Zanno, L. E., Gillette, D. D., Albright, L. B. & Titus, A. L., 2009: A new North American therizinosaurid and the role of herbivory in ‘predatory’ dinosaur evolution.
  –Proceedings of the Royal Society B: Biological Sciences: Vol. 276, #1672, pp. 3505-3511 [doi: 10.1098/rspb.2009.1029]