After Young & de Andrade, 2009, Young et al., 2012, and Parrilla-Bel et al., 2013, with taxa from Carroll, 1988, Clark, 1994, Storrs & Efimov, 2000, and Gasparini, Cichowolski & Lazo, 2005
<==o †Metriorhynchidae Fitzinger, 1843 [Metriorhynchi Fitzinger, 1843] |?- †Capelliniosuchus mutinensis Simonelli, 1896 [Capellineosuchus Romer, 1966, unjust. emend.?] [is a dolichosaurid mosasauroid lizard??] |--o †Geosaurinae Lydekker, 1889 sensu Young & Andrade, 2009 [Geosauridae Andrews, 1913] | |--o †Suchodus Lydekker, 1890 | | |-- †S. durobrivense Lydekker, 1890 [Metriorhynchus durobrivense (Lydekker, 1890)] | | `--+-- †S. cultridens (Andrews, 1913) [;Metriorhynchus cultridens Andrews, 1913] [=? Suchosaurus cultridens (Owen 1841) Owen, 1842, a spinosaurid tooth] | | `-- †S. brachyrhynchys (Deslongchamps, 1867) [Teleosaurus brachyrhynchus Deslongchamps, 1868; Metriorhynchus brachyrhynchys (Deslongchamps, 1867)] | `--+-- †Geosaurinae indet. [Argentina] | `--+--o †Purranisaurus Rusconi, 1948 | | |-- †P. potens Rusconi, 1948 [Metriorhynchus potens (Rusconi, 1948)] | | |-- †P. westermanni (Gasparini, 1980) [Metriorhynchus westermanni Gasparini, 1980] | | `-- †P. casamiquelai Gasparini & Chong, 1976 [Metriorhynchus casamiquelai Gasparini & Chong, 1976] | `--+-- †Neptunidragon ammoniticus Cau & Fanti, 2011 | `--+-- †Tyrannoneustes lythrodectikos Young, de Andrade, Brusatte, Sakamoto & Liston, 2013 | |-- †‘Mr Leeds’ specimen [GLAHM V972] | `--o †Geosaurini Lydekker, 1889 sensu Gau & Fanti, 2011 | |-- †Aggiosaurus nicaeensis (Ambayrac, 1913) Ambayrac, 1913 [Megalosaurus nicaeensis Ambayrac, 1913] | |-- †Plesiosuchus manselii (Hulke, 1870) Owen, 1884 [taxonomic history] | |-- †Torvoneustes carpenteri (Wilkinson, Young & Benton, 2008) Andrade, Young, Desojo & Brusatte, 2010 [Metriorhynchus superciliosus (de Blainville, 1853); Grange & Benton, 1996; Dakosaurus carpenteri Wilkinson, Young & Benton, 2008; Geosaurus carpenteri (Wilkinson, Young & Benton) Young & de Andrade, 2009] | |--o †Dakosaurus Quenstedt, 1852 [taxonomic history] | | |-- †D. andiniensis (Vignaud & Gasparini, 1996) [taxonomic history] | | `-- †D. maximus (Plieninger, 1846) [taxonomic history] | `--o †Geosaurus Cuvier, 1824 [taxonomic history] | |-- †G. lapparenti (Deblemas & Strannoloubsky, 1957) [Dakosaurus lapparenti Deblemas & Strannoloubsky, 1957] | `--+-- †G. grandis (Wagner, 1858) [Cricosaurus grandis Wagner, 1858; Cricosaurus medius Wagner, 1858] | `--+-- †G. sp. [Nusplingen Geosaurus, SMNS 81834] | `-- †G. giganteus(von Sömmerring, 1816) Cuvier, 1824 [taxonomic history] `--o †Metriorhynchinae Fitzinger, 1843 |--o †Metriorhynchus Meyer, 1832 [taxonomic history] | |?- †M. bathonicus Mercier, 1933 [taxonomic history] | |?- †M. blainvillei (Deslongchamps, 1866) | |?- †M. temporalis (Owen, 1884) | |--+-- †M. superciliosum (de Blainville, 1853) [taxonomic history] | | `-- †M. moreli Deslongchamps, 1869 | `--+-- †M. geoffroyi Meyer, 1832 [taxonomic history] | |?- †M. palpebrosus (Phillips, 1871) [jr. synon. M. geoffroyi?] | `--+-- †M. sp. [France] | `-- †M. hastifer (Deslongchamps, 1868) `--+--o †Gracilineustes Young et al., 2010 [Metriorhynchus (in partim)] | |-- †G. acutus Lennier, 1887 | `-- †G. leedsi (Andrews, 1913) `--+-- †Maledichosuchus riclaensis Parrilla-Bel, Young, Moreno-Azanza & Canudo, 2013 `--+-- †Metriorhynchidae indet. [Cuba] `--o †Rhacheosaurini |-- †Rhacheosaurus gracilis von Meyer, 1831 [Teleosaurus gracilis (Meyer); D'Alton & Burmeister, 1844; Geosaurus gracilis (Meyer, 1831)] |-- †Cricosaurus sp. [Cuba] `--o †Cricosaurus Wagner, 1858 [taxonomic history] |-- †C. saltillense (Buchy, Vignaud, Frey, Stinnesbeck & Gonzáles, 2006) |-- †C. suevicus (Fraas, 1902) [taxonomic history] |-- †C. elegans (Wagner, 1852) Wagner, 1958 [taxonomic history] `--+-- †C. vignaudi (Frey, Buchy, Stinnesbeck & Lopez-Oliva, 2002) [taxonomic history] |-- †C. gracilis (Philips, 1871) [taxonomic history] `--+-- †C. araucanensis (Gasparini & Dellapé, 1976) [taxonomic history] `--+-- †C. macrospondylus (Koken, 1889) [taxonomic history] `-- †C. schroderi (Kuhn, 1936) [taxonomic history]
Reference(s):
- de Andrade, M. B., Young, M. T., Desojo, J. B. & Brusatte, S. L., 2010: The Evolution of Extreme Hypercarnivory in Metriorhynchidae (Mesoeucrocodylia: Thalattosuchia) Based on Evidence from Microscopic Denticle Morphology.
–Journal of Vertebrate Paleontology: Vol. 30, #5, pp. 1451-1465 [doi: 10.1080/02724634.2010.501442] - Brochu, C. A. 1997: A review of “Leidyosuchus” (Crocodyliformes, Eusuchia) from the Cretaceous through Eocene of North America.
–Journal of Vertebrate Paleontology: vol. 17, #4, pp. 679-697 - Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698 - Carroll, R. L., 1988: Appendix. 594-648.
in Carroll, R. L. 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698 - Fernández, M. & Gasparini, Z., 2008: Salt glands in the Jurassic metriorhynchid Geosaurus: implications for the evolution of osmoregulation in Mesozoic marine crocodyliforms.
–Naturwissenschaften: Vol. 95, #1, pp. 79-84 [doi: 10.1007/s00114-007-0296-1] - Gasparini, Z., Cichowolski, M. & Lazo, D. G., 2005: First record of Metriorhynchus (Reptilia: Crocodyliformes) in the Bathonian (Middle Jurassic) of eastern Pacific.
–Journal of Paleontology: Vol. 79, #4, pp. 801-805 - Clark, J. M. 1994: Patterns of evolution in Mesozoic Crocodyliformes. 84-97
in Fraser, N. C. & Sues, H-D. 1994: In the Shadow of the Dinosaurs.
–Cambridge University Press, New York. 1994 - Pol, D. & Gasparini, Z., 2009: Skull anatomy of Dakosaurus andiniensis (Thalattosuchia: Crocodylomorpha) and the phylogenetic position of Thalattosuchia.
–Journal of Systematic Palaeontology: Vol. 7, #2, pp. 163-197 [doi: 10.1017/S1477201908002605] - Parrilla-Bel, J., Young, M. T., Moreno-Azanza, M. & Canudo, J. I., 2013: The First Metriorhynchid Crocodylomorph from the Middle Jurassic of Spain, with Implications for Evolution of the Subclade Rhacheosaurini.
–Plos One: Vol. 8, #1, pp. e54275 [doi: 10.1371/journal.pone.0054275] - Storrs, G. W. & Efimov, M. B., 2000: Mesozoic crocodyliforms of north-central Eurasia. 402-419
in Benton, M. J., Shishkin, M. A., Unvin, D. M. & Kurochkin, E. N., (eds.) 2000: The Age of Dinosaurs in Russia and Mongolia.
–Cambridge University Press, Cambridge, 2000 xxxix-696 - Young, M. T. & de Andrade, M. B., 2009: What is Geosaurus? Redescription of Geosaurus giganteus (Thalattosuchia: Metriorhynchidae) from the Upper Jurassic of Bayern, Germany.
–Zoological Journal of the Linnean Society: Vol. 157, #3, pp. 551-585 [doi: 10.1111/j.1096-3642.2009.00536.x] - Young, M. T., de Andrade, M. B., Brusatte, S. L., Sakamoto, M. & Liston, J., 2013: The oldest known metriorhynchid super-predator: a new genus and species from the Middle Jurassic of England, with implications for serration and mandibular evolution in predacious clades.
–Journal of Systematic Palaeontology: Vol. pp. 1-39 [doi: 10.1080/14772019.2012.704948] - Young, M. T., Brusatte, S. L., de Andrade, M. B., Desojo, J. B., Beatty, B. L., Steel, L., Fernández, M. S., Sakamoto, M., Ruiz-Omeñaca, J. I. & Schoch, R. R., 2012: The Cranial Osteology and Feeding Ecology of the Metriorhynchid Crocodylomorph Genera Dakosaurus and Plesiosuchus from the Late Jurassic of Europe.
–Plos One: Vol. 7, #9, pp. e44985 [doi: 10.1371/journal.pone.0044985]