After Holtz, 1998, and Smith, Makovicky, Hammer & Currie, 2007
<==o Tetanurae Gauthier, 1986 |--+-- †Condorraptor currumili Rauhut, 2005 | `-- †Piatnitzkysaurus floresi Bonaparte, 1979b `--+-- †Spinosauroidea Stromer, 1915 sensu Sereno et al., 1998 [Torvosauroidea, Megalosauria] `--o Neotetanurae Sereno |?-o †Megalosaurus Buckland, 1824 [paraphyletic? Megalosauridae Huxley, 1869] | |?- †M.? cambrensis (Newton, 1899) [Zanclodon cambrensis Newton, 1899] | |-- †M. hesperis Waldman, 1974 | `-- †M. bucklandii Ritgen, 1926 `--+--+?- †Streptospondylus altdorfensis Meyer, 1832 [†"Lealaps" gallicus Cope, 1867; Streptospondylus cuvieri (Owen, 1842) Huene, 1932; Eustreptospondylus divesensis Walker, 1964 partim] | |-- †Dubreuillosaurus valesdunensis (Allain, 2002) Allain, 2005 [†Poekilopleuron? valesdunensis Allain, 2002] | `-- †Eustreptospondylus oxoniensis Walker, 1964 [Eustreptospondylidae Paul, 1988] `--+--o †Torvosauridae | |?- †Edmarka rex Bakker, Krails, Siegwarth & Filla, 1992 | |?- †Magnosaurus nethercombensis (Huene, 1926b) [†"Megalosaurus" nethercombensis Huene, 1926b] | `-- †Torvosaurus tanneri Galton & Jensen, 1979a `--+-- †Afrovenator abakensi Sereno, Wilson, Larsson, Dutheil & Sues, 1994 `--o †Avetheropoda Paul, 1988 (avetheropods) |-- †Carnosauria [Allosauria Paul, 1988] (allosaurs and relatives; allosaurit) `-- Coelurosauria von Huene, 1914 sensu Holtz, 1999 (coelurosaurs; keelurosaurit) Tetanurae incertae sedis: †Becklespinax altispinax (Huene, 1923) sensu Olshevsky, 1991 [?Altispinax dunkeri (Dames, 1884), Megalosaurus dunkeri Dames, 1884, Acrocanthosaurus altispinax (Huene, 1923) sensu Paul, 1988] †“Brontoraptor” sp. Redman, 1995 [nomen nudum] †Cruxicheiros newmanorum Benson & Radley, 2010; M.Jur. WEu. †Kaijiangosaurus lini He, 1984 †Metriacanthosaurus parkeri Walker, 1964 †“Saltriosaurus” sp. Dal Sasso, 2000 [nomen nudum] †Sarcosaurus andrewsi Huene, 1932 [“Megalosaurus” andrewsi] †Sigilmassasaurus brevicollis Russell, 1996 †Unquillosaurus ceibalii Powell, 1979 †Xuanhanosaurus qilixiaensis Dong, 1984 †Chingkankousaurus fragilis Young, 1958a †Chuandongocoelurus primitivus He, 1984 †Iliosuchus incognitus Huene, 1932 [Iliosuchidae Paul, 1998] †Marshosaurus bicentesimus Madsen, 1976b †Kelmayisaurus petrolicus Dong, 1973
Note:
Phylogenetic position of so called “megalosaurs” is uncertain. These basal tetanurans have been often associated with spinosauroids (Smith, Makovicky, Hammer & Currie, 2007), but then again removed from that group. Here I have mainly followed Holtz (1998), by presenting a rather conservative view of basal Tetanurae.
My personal opinion is that megalosaurs are a paraphyletic basal group of Tetanurae and some of them are probably either basal avetheropods or basal spinosaurs.
Reference(s):
- Allain, R., 2002: Discovery of megalosaur (Dinosauria, Theropoda) in the middle Bathonian of Normandy (France) and its implications for the phylogeny of basal Tetanurae.
–Journal of Vertebrate Paleontology: Vol. 22, #3, pp. 548-563 - Allain, R., 2005: The postcranial anatomy of the megalosaur Dubreuillosaurus valesdunensis (Dinosauria Theropoda) from the Middle Jurassic of Normandy, France.
–Journal of Vertebrate Paleontology: Vol. 25, #4, pp. 850-858 - Benton, M. J., 1990: Origin and interrelationships of Dinosaurs. 11-30
in Weishampel, D. B., Dodson, P., & Osmólska, H. (eds.), 1990: The Dinosauria.
–University of California Press, Berkley, Los Angeles, Oxford, 1990 xvi-733 - Benson, R. B. J. & Radley, J. D., 2010: A new large-bodied theropod dinosaur from the Middle Jurassic of Warwickshirw, United Kingdom.
–Acta Palaeontologica Polonica: Vol. 55, #1, pp. 35-42 [doi: 10.4202/app.2009.0083] - Holtz, T. R. Jr., 1998 [preview]: Spinosaurs as crocodile mimics.
–Science: Vol. 282, 13 November, pp. 1276-1277 - Keesey, T. M., 1996-2000: Tetanurae – Three-fingered meat-eaters
–iNet: The Dinosauricon: http://dinosauricon.com/taxa/tetanurae.html - Novas, F. E. 1996: Dinosaur monophyly.
–Journal of Vertebrate Paleontology: vol. 16, #4, pp. 723-741. - Molnar, R., E., 1990: Problematic Theropoda: “Carnosaurs”. 306-317
in Weishampel, D. B., Dodson, P., & Osmólska, H. (eds.), 1990: The Dinosauria.
–University of California Press, Berkley, Los Angeles, Oxford, 1990 xvi-733 - Molnar, R. E., Kurzanov, S. M., & Dong, Z., 1990: Carnosauria. 169-209
in Weishampel, D. B., Dodson, P., & Osmólska, H. (eds.), 1990: The Dinosauria.
–University of California Press, Berkley, Los Angeles, Oxford, 1990 xvi-733 - Padian, K., Hutchinson, J. R. & Holtz, T. R. Jr., 1999: Phylogenetic defenitions and nomenclature of the major taxonomic categories of the carnivorous Dinosauria (Theropoda).
–Journal of Vertebrate Paleontology: Vol. 19, #1, pp. 69-80 - Paul, G. S. 1988: The Predatory Dinosaurs of the World.
–Simon and Schuster Co., New York, 1988. 464. - Rauhut, O. W. M., 2005: Osteology and relationships of a new theropod dinosaur from the Middle Jurassic of Patagonia.
–Palaeontology: Vol. 48, #1, pp. 87-111