After Hungerbühler, 2002
<==o †Phytosauria Jaeger, 1828, non v. Meyer, 1861 [Parasuchia Huxley, 1875] |?-o †Paleorhinus | |--+-- †P. bransoni | | `-- †P. sawini | |-- †P. fraasi | |-- †P. ehlersi | |-- †P. magnoculus | `-- †P. scurriensis `--o †Phytosauridae Jaeger, 1828, non v. Meyer, 1861 sensu Hungerbühler, 2002 |?- †Belodon plieningeri von Meyer, in von Meyer & Plieningei, 1844 |?-o †Clepsysaurus Lea, 1851 [a phytosaur?] | |-- †C. pennsylvanicus Lea, 1851 [tooth-taxa] | `-- †C. veatleianus [tooth-taxa] |?- †Ebrachosuchus neukami Kuhn, 1936 [Paleorhinus neukami (Kuhn, 1936)] |?- †Parasuchus hislopi |?- †Mesorhinosaurus fraasi |--o †Rutiodon Emmons, 1856 [Angistorhinus; Machaeroprosopus Mehl, 1916] | |-- †R. grandis [Angistorhinus grandis] | |-- †R. megalodon [Brachysuchus megalodon] | |-- †R. carolinensis Emmons, 1856 | |-- †R. sp. | |-- †R. crosbiensis | |-- †R. gregorii [=? Rutiodon gregoryi?; Smilosuchus gregorii?; Machaeroprosopus gregorii; Nicrosaurus gregoryi; Redondasaurus gregorii] | |-- †R. lepturus | `-- †R. manhattanensis Huene, 1913 |-- †Smilosuchus gregorii [=? Rutiodon gregoryi?] |--o †Leptosuchus Case, 1922 | |-- †L. crosbiensis Case, 1922 [Phytosaurus doughtyi Case, 1920] | |-- †L. studeri Case & White, 1934 | |-- †L. talainti [Angistorhinus talainti; Rutiodon talainti] | `-- †L. adamanensis [Rutiodon adamanensis] |?-o †Phytosaurus | |?- †P. cylindricodon Jaeger, 1828 [nomen dubium] | |-- †P. doughtyi | `-- †P. scolopax `--o †Pseudopalatinae Long & Murry, 1995 [incl. Mystriosuchinae v. Huene, 1915] |?- †Angistorhinopsis ruetimeyeri (Huene, 1922) |?- †Coburgosuchus goeckili Heller, 1954 |--o †Nicrosaurus O. Fraas, 1866 [Phytosaurus] | |-- †N. kapffi (von Meyer, 1860) [Phytosaurus kapffi von Meyer, 1860] [type] | |-- †N. gregoryi [=? Rutiodon gregoryi?; Smilosuchus gregorii?] | `-- †N. meyeri Hungerbühler & Hunt, 2000 `--+--o †Mystriosuchus E. Fraas, 1896 [taxonomic history] | |-- †M. planirostris (von Meyer, 1863) [taxonomic history] | `-- †M. westphali Hungerbühler & Hunt, 2000 [taxonomic history] `--+--o †Redondasaurus | |-- †R. gregorii [=? Rutiodon gregoryi?; Smilosuchus gregorii?; Nicrosaurus gregoryi?] | `-- †R. bermani `--o †Pseudopalatus |?- †P. andersoni |-- †P. mccauleyi `--+-- †P. pristinus `-- †P. buceros [Phytosaurus buceros; Nicrosaurus buceros; Arribasuchus buceros]
Phytosauria incertae sedis:
†Palaeoctonus Cope, 1877 †P. appalachianus [tooth-taxa] †P. aulacodus [tooth-taxa]
Reference(s):
- Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698 - Carroll, R. L., 1988: Appendix. 594-648.
in Carroll, R. L. 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698. - Doyle, K. D. & Sues, H.-D., 1995: Phytosaurs (Reptilia: Archosauria) from the Upper Triassic New Oxford Formation of York County, Pennsylvania.
–Journal of Vertebrate Paleontology: Vol. 15, #3, pp. 545-553 - Hungerbühler, A., 2001: Status and phylogenetic relationships of the Late Triassic phytosaur Rutiodon carolinensis.
–Journal of Vertebrate Paleontology: Vol. 21, #3, Supplement to #3, pp. 64A - Hungerbühler, A., 2002: The Late Triassic phytosaur Mystriosuchus westphali, with a revision of the genus.
–Palaeontology: Vol. 45, #2, pp. 377-418 - Parrish, J. M. 1993: Phylogeny of the Crocodylotarsi, with Reference to Archosaurian and Crurotarsan Monophyly.
–Journal of Vertebrate Paleontology: vol. 13, #3, pp. 287-308 - Sereno, P. C. 1991: Basal Archosaurs: phylogenetic relationships and functional implications.
–Journal of Vertebrate Paleontology: Vol. 11, Supplement to #4, Memoir 2. pp. ii-53
Special thanks to:
Jos H. M. Dols, for gathering additional information and species.