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After Wagner, 2002
<==o †Euomphalina de Koninck, 1881 |-- †Prohelicotoma |-- †Ophileta subraplana `--+-- †Ophileta complanata `--+-- †Lecanospira compacta |-- †Lecanospira nereine `--+-- †Barnesella ?lecanospiroides |--o †Ecculiomphalus Portlock, 1843 | |-- †Malayaspira rugosa | `--+-- †MACLURITOIDEA Fischer, 1885 | `--+-- †Malayaspira hintzei | |--+--+-- †Russospira harrisae | | | `-- †Ecculiomphalus bucklandi | | `--o †Helicotomidae Wenz, 1938 | | |?- Linsleyella Rohr, 1980 | | |-- †Lophonema peccatonica Ulrich, in Purdue & Miser, 1916 | | |--+-- Polehemia taneyensis | | | `-- Walcottoma frydai | | `--+--+--o Ophiletina Ulrich & Scofield, 1897 | | | | |-- Oriostoma bromidensis | | | | `--+-- O. sublaxa | | | | `-- O. angularis | | | `--o †Helicotoma Salter, 1859 | | | |-- H. tennesseensis | | | |--+-- H. planulata | | | | `--+-- H. robinsoni | | | | `-- H. blodgetti | | | `--o Palaeomphalus Koken, 1925 | | | |-- P. ?gradatus | | | `-- Helicotoma? sp. [Girvan sp.] | | `--+--o Anomphalidae Wenz, 1938 | | | `--o †Trochomphalus Koken, 1925 | | | |-- Trochomphalus ?dimidiatus | | | `--+-- Straporillina sp. cf. S. circe (sensu Rohr, 1988) | | | `--+-- Grantlandispira christei | | | `--o Pycnomphalus Lindström, 1884 | | | |-- P. acutus | | | `--+-- P. obesus | | | `-- Turbocheilus immaturum | | `--o †Euomphalidae de Koninck, 1881 | | |?- Micromphalus Knight, 1945 | | |--o †Boucotspira Rohr, 1980 | | | |-- B. cariniferus [Euomphalopterus cariniferus] | | | `-- B. ?ordovicicus [Euomphalopterus ?ordovicicus] | | |--o †Euomphalopterus Roemer, 1876 | | | |-- E. subcarinatus | | | |-- E. alatus | | | `--+-- E. togatus | | | `--+--+-- E. praetextus | | | | `-- E. undulans | | | `--o Spinicharybdis Rohr & Packard, 1982 | | | |-- S. frenatus [Euomphalopterus frenatus] | | | `-- S. wilsoni | | |--o †Poleumita Clarke & Ruedemann, 1903 | | | |-- P. alata | | | `--+-- P. discors | | | |-- P. granulosa | | | |-- Centrifugus planorbis Bronn, 1834 | | | |--o Nodonema Linsley, 1968 | | | | |-- N. rugosa [Poleumita rugosa] | | | | `-- N. octavia [Poleumita octavia] | | | `--+-- Euomphalus Sowerby, 1814 | | | `-- Straparollus de Montfort, 1810 | | `--o Pseudophoridae Miller, 1889 | | |?- Elasmonema Fischer, 1885 | | |-- Streptotrochus lamellosus [Boucotspira [in partim]] | | `--+-- Streptotrochus? visbeyensis [Boucotspira [in partim]] | | `--+--o Streptotrochus Perner, 1903 | | | |-- S. incisus | | | `--+-- S. lundgreni | | | `-- Hystricoceras astraciformis Jahn, 1894 | | `--o †Discordichilus Cossmann, 1918 | | |-- Streptotrochus sp. aff. S. incisus | | `--+-- D. gotlandicus [Siluriphorus gotlandicus] | | |-- D. undulans [Siluriphorus undulans] | | |--+--+-- D. dalli | | | | `-- D. mollis | | | `--o Pseudotectus Perner, 1903 | | | |-- P. kolmodoni [Discordichilus kolmodoni] | | | `-- P. comes | | `--o Pseudophorus Meek, 1873 | | |-- P. stuxbergi | | `-- P. profundus | `--o Lesueurillidae Wagner, 2002 | |-- †"Lesueurilla" declivis | |--o †Eccyliopterus Remelé, 1888 | | |-- E. regularis | | `--+-- E. ?princeps | | `--+-- E. alatus | | `-- E. owenanus | `--+-- "Lesueurilla" prima | |--o †Lesueurilla Koken, 1898 | | |-- "Eccyliopterus" louderbacki | | `--+-- Lesueurilla infundibula | | `-- "Eccyliopterus" beloitensis | |--o Mestoronema Wagner, 2002 | | |-- "Lesueurilla" marginalis | | `--+-- "Lesueurilla" scotica | | `-- "Lesueurilla" bipatellare | `--o Pararaphistoma Vostokova, 1955 | |-- Climacoraphistoma vaginati | |-- Climacoraphistoma damesi | `--+-- Pararaphistoma qualteriana | `-- Pararaphistoma schmidti `--o †Asgardaspira Wagner, 2002 |-- †Barnesella measuresae `--+-- †Lytospira yochelsoni `--+--o †Lytospira Koken, 1896 | |-- †L. angelini | `-- †L. subrotunda `--+--+-- †Lytospira gerrula | `-- †Lytospira ?norvegica `--o †Euomphaloidea de Koninck, 1881 `--o †Raphistomatidae Koken, 1896 |-- †Ceratopea canadensis |-- Orospira bigranosa Butts, 1926 |--o Bridgeites Flower, 1968a | |-- †B. planodorsalis | `--+-- †B. supraconvexa | `-- †B. ?disjuncta `--+-- †Ceratopea ?laurentia |--+-- †Ceratopea pygmaea | `-- †Ceratopea unguis ==> `--+-- Pararaphisoma lemoni `--+-- †Helicotoma medffraensis Salter, 1859 `--+--o †Scalites Emmons, 1842 | |-- “Palaeomphalus” giganetus Koken, 1925 ==> | `--+-- Scalites katoi | `--+--+-- Raphistoma tellerensis | | `-- Raphistoma peracuta | `--+-- Scalites angulatus | `--o Raphistoma Hall, 1847 | |-- Raphistoma striata | `-- Helicotoma gubanovi `--o Holopeidae Wenz, 1938 |?- Umbospira Perner, 1903 |--o Raphistomina Ulrich & Scofield, 1897 | |-- R. lapicida | |-- R. aperta | |-- R. fissurata | `-- R. rugata `--+-- Pachystrophia devexa |--+-- Pachystrophia contigua | `--o Holopea Hall, 1847 | |-- H. insignis | |-- H. pyrene | `--+-- H. rotunda | `-- H. symmetrica `--+--o Sinutropis Perner, 1903 | |-- Pachystrophia spiralis | |-- Sinutropis ?esthetica | `-- Euomphalus tubus `--o †Pachystrophia Perner, 1903 |-- P. gotlandica `--+-- Lytospira triquestra `-- Lytospira subuloides Euomphalina de Koninck, 1881 [fide Wagner, n.d. & Nützel, 3/2002] [Euomphalomorpha Bandel & Frýda, 1998 (Nützel, 3/2002); Macluritina Cox & Knight, 1960] Ophiletoidea Knight, 1956 Ophiletidae Knight, 1956 Macluritoidea P. Fischer, 1885 ?Archaeospiridae Yu, 1979 (Lower Cambrian) Macluritidae P. Fischer, 1885 [Macluridae Carpenter, 1858; Macluriidae Pilsbry, 1888] Euomphaloidea deKoninck, 1881 [Omphalotrochacea Knight, 1945; Anomphaloidea Wenz, 1938; Pseudophoroidea Miller, 1889 fide Wagner; “Rhaphistomatoidea” Koken, 1896 sic of Golikov & Starobogatov, 1975] Anomphalidae Wenz, 1938 Euomphalidae de Koninck, 1881 [Schizostomidae Bronn, 1849 (& Eichwald, 1871); Polytropidae Koken, 1925; Poleumitidae Wenz, 1938] Helicotomidae Wenz, 1938 Holopeidae Wenz, 1938 Lesueurillidae Wagner, 2002 Omphalotrochidae Knight, 1945 Pseudophoridae Miller, 1889 [Palaeonustidae Wenz, 1938] Planitrochidae Knight, 1956 Raphistomatidae Koken, 1896 [Raphistomidae Koken, 1896] Platyceratoidea Hall, 1859 Platyceratidae Hall, 1859 [Platyceridae Hall, 1859; Platystomidae Miller, 1889; Cyclonemidae Miller, 1889]
Note: Phylogenetic diagrams of Wagner (2002) were particularly difficult to understand, mainly because his analysis was so huge, that he had to split the result tree to several subtrees, which still inclueded several other subtrees. I hope I got it right. :-(
Wagner, P. J., 2002 [1995]: Phylogenetic relationships of the earliest anisostrophically coiled gastropods.
--Smithsonian Contributions to Paleobiology: Vol. 88, pp. vi-152
[Author [Wagner] notes that scientific material of this article is based on his 1995 Ph.D. -thesis, which is partly outdated]
© Mikko Haaramo, / Last updated 2003-02-21 / http://www.mv.helsinki.fi/home/mhaaramo