After Parker, 1982
<==o Tardigrada (karhukaiset; water bears) |--o Heterotardigrada | |--o Arthrotardigrada | | |-- Halechiniscidae | | |-- Batillipedidae | | `-- Stygarctidae | `--o Echiniscoidea | |-- Oreellidae | `-- Echiniscidae `--+--o Mesotardigrada | `-- Thermozodiidae `--o Eutardigrada |-- Macrobiotidae `-- Milnesiidae
After ITIS
<==o Tardigrada (Spallanzani, 1777) (karhukaiset; water bears) |--o Heterotardigrada Marcus, 1927 | |--o Arthrotardigrada Marcus, 1927 | | |--o Batillipedidae Ramazzotti, 1962 | | | `--o Batillipes Richters, 1909 | | |--o Coronarctidae Renaud-Mornant, 1974 | | | |--o Coronarctus Renaud-Mornant, 1974 | | | `--o Trogloarctus Villora-Moreno, 1996 | | |-- Halechiniscidae Thulin, 1928 | | |--o Neostygarctidae Grimaldi de, d’Addabbo Gallo & Moreno de Lucia, 1987 | | | `--o Neostygarctus Grimaldi de, d’Addabbo Gallo & Moreno de Lucia, 1982 | | |--o Renaudarctidae Kristensen & Higgings, 1984 | | | `--o Renaudarctus Kristensen & Higgings, 1984 | | `--o Stygarctidae Schulz, 1951 | | |--o Megastygarctinae Bello & Grinmaldi de Zio, 1999 | | | `--o Megastygartides McKirdy, Schmidt & McGinty-Bayly, 1976 | | |--o Stygarctinae Schulz, 1951 | | | |--o Mesostygarctus Renaud-Mornant, 1979 | | | |--o Parastygarctus Renaud-Debyser, 1965 | | | |--o Pseudostygarctus McKirdy, Schmidt & McGinty-Bayly, 1976 | | | `--o Stygarctus Schulz, 1951 | | `--o Neoarctinae Grimaldi de, d’Addabbo Gallo & Moreno de Lucia, 1992 [= Neoartctidae Grimaldi de, d’Addabbo Gallo & Moreno de Lucia, 1992] | | `--o Neoarctus Grimaldi de, d’Addabbo Gallo & Moreno de Lucia, 1992 | `--o Echiniscoidea Marcus, 1927 | |--o Carphaniidae Binda & Kristensen, 1986 | | `--o Carphania Binda, 1979 | |--o Echiniscidae Thulin, 1928 | | |--o Antechiniscus Kristensen, 1987 | | |--o Bryochoerus Marcus, 1936 | | |--o Bryodelphax Thulin, 1928 | | |--o Cornechiniscus Maucci & Ramazzotti, 1981 | | |--o Echiniscus C.A.S. Schultze, 1840 | | |--o Hypechiniscus Thulin, 1928 | | |--o Mopsechiniscus Du Bois-reymond Marcus, 1944 | | |--o Novechiniscus Kristensen, 1987 | | |--o Parechiniscus Cuénot, 1926 | | |--o Proechiniscus Kristensen, 1987 | | |--o Pseudechiniscus Thulin, 1911 | | `--o Testechiniscus Kristensen, 1987 | |--o Echiniscoididae Kristensen & Hallas, 1980 | | |--o Anisonyches Pollock, 1975 | | `--o Echiniscoides Plate, 1889 | `--o Oreellidae Puglia, 1959 | `--o Oreella Murray, 1910 `--+--o Mesotardigrada Rahm, 1937 | `--o Thermozidia Ramazzotti & Maucci, 1983 | `--o Thermozodiidae Rahm, 1937 | `--o Thermozodium Rahm, 1937 `--o Eutardigrada Richters, 1926 |--o Apopchela Schuster, Nelson, Grigarick & Christenberry, 1980 | `--o Milnesiidae Ramazzotti, 1962 | |--o Limmenius Horning, Schuster & Grigarick, 1978 | |--o Milnesioides Claxton, 1999 | `--o Milnesium Doyère, 1840 `--o Parachela Schuster, Nelson, Grigarick & Christenberry, 1980 |?-o Beornidae Cooper, 1964 [? incl. in Hybsibiidae Pilato, 1969] | `--o Beorn Cooper, 1964 |--o Calohybsibiidae Pilato, 1969 | |--o Calohypsibius Thulin, 1928 | |--o Haplohexapodibius Pilato & Beasley, 1987 | |--o Haplomacrobiotus May, 1948 | |--o Hexapodibius Pilato, 1969 | `--o Parhexapodibius Pilato, 1969 |--o Eohypsibiidae Bertolani & Kristensen, 1987 | |--o Amphibolus Bertolani, 1981 | `--o Eohypsibius Kristensen, 1982 |-- Hybsibiidae Pilato, 1969 |--o Macrobiotidae Thulin, 1928 | |--o Macrobiotinae Guidetti, Rebecchi & Bertolani, 2000 | | |--o Adorybiotus Maucci & Ramzzotti, 1981 | | |--o Biserovus Guidetti & Pilato, 2003 | | |--o Calcarobiotus Dastych, 1993 | | |--o Famelobiotus Pilato, Binda & Lisi, 2004 | | |--o Insuetifurca Guidetti & Pilato, 2003 | | |--o Macrobiotus Schultze, 1834 | | |--o Minibiotus Schuster, 1980 | | |--o Minilentus Guidetti & Pilato, 2003 | | |--o Pseudodiphascon Ramazzotti, 1965 | | |--o Pseudohexapodibius Bertolani & Biserov, 1996 | | |--o Richtersius Pilato & Binda, 1990 | | `--o Xerobiotus Bertolani & Biserov, 1996 | `--o Murrayinae Guidetti, Rebecchi & Bertolani, 2000 | |--o Dactylobiotus Schuster, 1980 | |--o Macroversum Pilato & Catanzaro, 1988 | `--o Murrayon Bertolani & Pilato, 1989 |--o Microhybsibiidae Pilato, 1998 | |--o Fractonotus Pilato, 1998 | `--o Microhypsibius Thulin, 1928 `--o Necopinatidae Ramazzotti & Maucci, 1983 |--o Apodibius Dastych, 1983 `--o Necopinatum Pilato, 1971
Reference(s):
–iNet: http://www.itis.gov/servlet/SingleRpt/SingleRpt?search_topic=TSN&search_value=155359- Parker, S. P. (ed.), 1982: Synopsis and classification of living organisms. Vols. 1 & 2
–McGrew-Hill Book Company
Special thanks to:
Jos Dos, for gathering information about the species and genera.