After Grimaldi & Engel, 2005 (based on Zwick, 2000)
<==o Plecoptera Latreille, 1902 sensu lato [incl.? †“Protoperlaria”] (koskikorennot; stoneflies) |--o †Lemmatophoridae Sellards 1909 [Ortaidae Sellards, 1909; Lecoriidae Handlirsch, 1919] [paraphyletic †“Protoperlaria”] | |--o †Lemmatophora Sellards 1909 | | `-- †T. typa Sellards 1909 [T. typica Tillyard 1928; T. anomala; T. elongata; T. delicosa; T. hirsuta, Sellards, 1909] | |--o †Artinska Sellards 1909 [Estadia Sellards, 1909; Lectrum Sellards, 1909; Orta Sellards, 1909] | | `-- †A. clara Sellards 1909 [A. medialis Sellards, 1909; A. gracilis Sellards, 1909; A. pecta Sellards, 1909; A. major Sellards, 1909; A. extensa Sellards, 1909; Estadia elongata Sellards, 1909;, E. arcuata Sellards, 1909; E. tenuis Sellards, 1909; Estadia tripunctata Carpenter 1926; Lectrum anomalum Sellards, 1909] | `--o †Lisca Sellards, 1909 | `-- †L. minuta Sellards, 1909 [Lemmatophora reducta Tillyard 1928; L. minuta Tillyard 1928; L. cubitalis Handlirsch 1937] |--o †Liomopteridae Sellards, 1909 [paraphyletic †“Protoperlaria”] | `--o †Liomopterum Sellards 1909 [Horates Sellards, 1909] | `-- †L. ornatum Sellards, 1909 [L. extensum Sellards, 1909] `--+--o †Probnidae Sellards, 1909 [nom. correct. pro Probnisidae Sellards 1909] | `--o †Probnis Sellards 1909 [= Espira Sellards 1909; Stoichus Sellards 1909; Stinus Sellards 1909] | `-- †P. speciosa Tillyard 1937 [P. coriacea Sellards, 1909; Espira obscura Sellards, 1909; Stoichus elegans Sellards, 1909; S. arcuatus Sellards, 1909; S. minor Sellards, 1909; S. tenuis Sellards, 1909; Stinus brevi-cubitalis Sellards 1909; Probnis sellardsi Handlirsch 1919] `--o Plecoptera sensu Grimaldi & Engel, 2005 |-- †Palaeonemouridae |-- †Palaeoperlidae |-- †Perlopseidae `--+--o Antarctoperlaria sensu lato | |-- †“Eusthenioides” | |-- †Euxenoperlidae | `--+?- †Siberioperlidae Sinitshenkova, 1983 | `--o Antarctoperlaria | |--o Eusthenioidea | | |-- Eustheniidae | | `-- Diamphipnoidae | `--o Leptoperloidea [Griphopterygoidea] | |-- Austroperlidae | `-- Gripopterygidae `--o Arctoperlaria |--o Systellognatha [Subulipalpia]? | |?- †Platyperlidae | `--+--+-- Pteronarcyidae | | `--+-- Peltoperlidae | | `--o Styloperlidae | | |-- Styloperla | | `-- Cerconychia | `--+-- Perlodidae | |-- Perlidae | `-- Chloroperlidae `--o Euholognatha |-- Scopuridae `--o Nemouroidea? |?- †Perlariopseidae |-- Taeniopterygidae `--+?- †Baleyopterygidae |--+-- “Notonemouridae” | `-- Nemouridae `--+-- Capniidae `-- Leuctridae
Alternative phylogeny after Parker, 1982, and Nelson, 1996-1998 [“The Tree of Life”]
<==o Plecoptera [incl.? †“Protoperlaria”] (koskikorennot; stoneflies) |--o Eusthenioidea | |-- Eustheniidae | `-- Diamphipnoidae `--+--o Leptoperloidea | |-- Austroperlidae | `-- Gripopterygidae `--o EUHOLOGNATHA? [paraphyletic?] |-- Scopuridae |--o NEMOUROIDEA? | |-- Taeniopterygidae | `--+--+-- Notonemouridae | | `-- Nemouridae | `--+-- Capniidae | `-- Leuctridae `--o SYSTELLOGNATHA [Subulipalpia]? |-- Pteronarcyidae `--+--+-- Peltoperlidae | `--o Styloperlidae | |-- Styloperla | `-- Cerconychia `--+-- Perlodidae |-- Perlidae `-- Chloroperlidae
Reference(s):
- Beckemeyer, R. J. 2006-: Checklist of Midco, Oklahoma Permian insects. [visited: 2006-08-04]
–iNet: www.windofkansas.com: http://www.windsofkansas.com/midco_taxa.html - Grimaldi, D. A. & Engel, M. S., 2005: Evolution of the Insects.
–Cambridge University Press, New York, 2005, xv-755 - Nelson, C. R., 1996: Styloperlidae.
–iNet: The Tree of Life Project: http://www.utexas.edu/courses/capnia/plecoptera/styloperlidae/styloperlidae.html - Nelson, C. R., 1996-1998: Plecoptera – Stoneflies.
–iNet: The Tree of Life Project: http://www.utexas.edu/courses/capnia/plecoptera/plecoptera.html - Parker, S. P. (ed.), 1982: Synopsis and classification of living organisms. Vols. 1 & 2
–McGrew-Hill Book Company