After Tollerton, 1989

<==o Eurypterida Burmeister, 1843 (meriskorpionit; seascorpions)
   |--o †Eurypterina Burmeister, 1843 (nom. trans. Caster & Kjellesvig-Waering, 1964)
   |  |--o †Slimonioidea Novojilov, 1962
   |  |  `--o Slimoniidae Novojilov, 1962
   |  |     |-- †Slimonia Page, 1856
   |  |     `-- †Salteropterus Kjellesvig-Waering, 1951
   |  |--o †Hughmillerioidea Kjellesvig-Waering, 1951 [nom. trans. Størmer, 1974]
   |  |  |--o †Hughmilleriidae Kjellesvig-Waering, 1951
   |  |  |  |-- †Hughmilleria Sarle, 1903
   |  |  |  |-- †Nanahughmilleria Kjellesvig-Waering, 1961b
   |  |  |  |-- †Grossopterus Størmer, 1934b
   |  |  |  `-- †Waeringopterus Leutze, 1961
   |  |  |--o †Carcinosomatidae Størmer, 1934 [Carcinosomidae Størmer, 1934]
   |  |  |  |?- †Holmipterus Kjellesvig-Waering, 1979
   |  |  |  |-- †Carcinosoma Claypole, 1890 [pro Eurysoma Claypole, 1890 (non Koch, 1839; nec Dujardin, 1845)]
   |  |  |  |-- †Eocarcinosoma Caster & Kjellesvig-Waering, 1964
   |  |  |  |-- †Paracarcinosoma Caster & Kjellesvig-Waering, 1964
   |  |  |  `-- †Rhinocarcinosoma Novojilov, 1962
   |  |  `--o †Adelophthalmidae Tollerton, 1989
   |  |     |-- †Adelophthalmus Jordan & Meyer, 1854
   |  |     |-- †Parahughmilleria Störmer, 1974
   |  |     |-- †Bassipterus Kjellesvig-Waering & Leutze, 1966
   |  |     `-- †Unionopterus Chernyshev, 1948
   |  |--o †Mixopteroidea Caster & Kjellesvig-Waering, 1955
   |  |  |--o †Mixopteridae Caster & Kjellesvig-Waering, 1955)
   |  |  |  `-- †Mixopterus Ruedemann, 1921
   |  |  `--o †Lanarkopteridae Tollerton, 1989
   |  |     `-- †Lanarkopterus Ritchie, 1968
   |  |--o †Megalograptoidea Caster & Kjellesvig-Waering, 1955 [nom. trans. Tollerton, 1989]
   |  |  `--o †Megalograptidae Caster & Kjellesvig-Waering, 1955
   |  |     |-- †Echinognathus Walcott, 1882
   |  |     `-- †Megalograptus Miller, 1874
   |  |--o †Eurypteroidea Burmeister, 1843 [Eurypteraceae Burmeister, 1843]
   |  |  |--o †Eurypteridae Burmeister, 1843
   |  |  |  |?- †Onychopterella Størmer, 1951 [pro Onychopterus Clarke & Ruedemann, 1912 (non Reichenbach, 1850); Eurypterus kokomoensis Miller & Gurley, 1896]
   |  |  |  `-- †Eurypterus De Key, 1825 [non Mabille, 1877; nec Sharpe, 1896] [incl. Baltoeurypterus Størmer, 1973 sensu Tetlie, 2006]
   |  |  |--o †Dolichopteridae Kjellesvig-Waering & Størmer, 1952
   |  |  |  |-- †Dolichopterus hall, 1859
   |  |  |  |-- †Strobilopterus Ruedemann, 1935a
   |  |  |  |-- †Ruedemannipterus Kjellesvig-Waering, 1966
   |  |  |  `-- †Syntomopterus Kjellesvig-Waering, 1961a
   |  |  `--o †Erieopteridae Tollerton, 1989
   |  |     |-- †Buffalopterus Kjellesvig-Waering & Heubusch, 1962
   |  |     |-- †Onychopterella Størmer, 1951 [pro Onychopterus Clarke & Ruedemann, 1912 (non Reichenbach, 1850)]
   |  |     `-- †Erieopterus Kjellesvig-Waering, 1958
   |  |--o †Stylonuroidea Diener, 1924 [Stylonauracea Diener, 1924; Drepanopteroidea Kjellesvig-Waering, 1966]
   |  |  |--o †Stylonuridae Diener, 1924 [Pageidae Kjellesvig-Waering, 1966, in Waterston, 1979]
   |  |  |  |-- †Stylonurus Page, 1856
   |  |  |  |-- †Stylonurella Kjellesvig-Waering, 1966
   |  |  |  `-- †Kiaeropterus Waterston, 1979
   |  |  |--o †Drepanopteridae Kjellesvig-Waering, 1966
   |  |  |  |-- †Drepanopterus Laurie, 1892
   |  |  |  |-- †Alkenopterus Størmer, 1974
   |  |  |  `-- †Moselopterus Størmer, 1974
   |  |  |--o †Parastylonuridae Waterston, 1979
   |  |  |  |?- ?†Stylonuroides Kjellesvig-Waering, 1966
   |  |  |  |?- ?†Brachyopterus Størmer, 1951
   |  |  |  `-- †Parastylonurus Kjellesvig-Waering, 1966
   |  |  `--o †Laurieipteridae Kjellesvig-Waering, 1966
   |  |     |-- †Laurieipterus Kjellesvig-Waering, 1966
   |  |     |-- †Ctenopterus Clark & Ruedemann, 1912
   |  |     |-- †Hallipterus Kjellesvig-Waering, 1963a
   |  |     |-- †Pagea Waterston, 1962
   |  |     `?- ?†Mazonipterus Kjellesvig-Waering, 1963b
   |  |--o †Kokomopteroidea Kjellesvig-Waering, 1966 [nom. trans. Tollerton, 1989]
   |  |  |--o †Kokomopteridae Kjellesvig-Waering, 1966
   |  |  |  |-- †Kokomopterus Kjellesvig-Waering, 1966
   |  |  |  `-- †Lamontopterus Waterston, 1979
   |  |  `--o †Hardieopteridae Tollerton, 1989
   |  |     `-- †Hardieopterus Waterston, 1979
   |  |--o †Brachyopterelloidea Tollerton, 1989
   |  |  `--o †Brachyopterellidae Tollerton, 1989
   |  |     `-- †Brachypterella Kjellesvig-Waering, 1966
   |  `--o †Mycteropoidea Cope, 1886
   |     |--o †Mycteropidae Cope, 1886
   |     |  `--o †Mycterops Cope, 1886 [Glaucodes Pruvost, 1923]
   |     |     |-- †M. scabrosus (Woodward, H.)
   |     |     `-- †M. mathieui (Pruvost, 1923) [Glaucodes mathieui Pruvost, 1923]
   |     `--o †Woodwardopteridae Kjellesvig-Waering, 1959
   |        |-- †Woodwardopterus Kjellesvig-Waering, 1959
   |        `-- †Vernopterus Waterson, 1968
   `--o †Pterygotina Caster & Kjellesvig-Waering, 1964
      `--o †Pterygotoidea Clarke & Ruedemann, 1912
         |--o †Jaekelopteridae Størmer, 1974
         |  `-- †Jaekelopterus Waterston, 1964
         `--o †Pterygotidae Clarke & Ruedemann, 1912
            |-- †Pterygotus Agassiz, 1839
            |-- †Acutiramus Ruedemann, 1935b
            |-- †Erettopterus Salter, 1859 [pro Himantopterus Salter, 1856 (non Westmail, 1836)]
            `-- †Truncatiramus Kjellesvig-Waering, 1961b

Eurypterida incertae sedis:

    †Melbournopterus Caster & Kjellesvig-Waering, 1953
    †Dorfopterus Kjellesvig-Waering, 1955
    †Hastimima White, 1908
    †Pittsfordipterus Kjellesvig-Waering & Leutze, 1966
    †Tarsopterella Størmer, 1951 [pro Tarsopterus Clarke & Ruedemann, 1912 (non Reinhardt & Luetken, 1862)]
    †Tylopterella Størmer, 1951 [pro Tylopterus Clarke & Ruedemann, 1912 (non Capiomont, 1868; nec Leconte, 1876)]
    †Willwerathia Størmer, 1969
    ?†Campulocephalus Eichw., 1860 [not recognized by Tollerton (1989)]
    ?†Lepidoderma Reuss, 1855 [non Waterhouse, 1875, nec Zlinka, 1889] [not recognized by Tollerton (1989)]
    ?†Glyptoscorpius Peach, 1882 [not recognized by Tollerton (1989)]
    ?†Belinuropsis Matthew, 1909 [not recognized by Tollerton (1989)]

Reference(s):