After Ahyong, Lai, Sharkey, Colgan & Ng, 2007, Parker, 1982, and Martin & Davis, 2001; with Recent genera and species from the “WoRMS – World Record of Marine Species”
Alternative, older, taxonomy after Parker, 1982
<==o Brachyura Latreille, 1802 (crabs; taskuravut) |-- Dromiacea de Haan, 1833 `--+--o Raninoidea de Haan, 1839 | |--o Symethis Weber, 1795 [non Symenthis Goeke] [Symethidae Goeke, 1981] | | |-- S. corallica Davie, 1989 | | |-- S. garthi Goeke, 1981 | | `-- S. variolosa (Fabricius, 1793) | `--o Raninidae de Haan, 1839 | |-- †Erroranilia Boyko, 2004 [†Paralbunea Hu & Tao, 1996] | |-- Cyrtorhina Monod, 1956 [Cyrtorhininae Guinot, 1993] | |--o Lyreidinae Guinot, 1993 | | |-- Lyreidus De Haan, 1841 | | `-- Lysirude Goeke, 1985 | |--o Notopodinae Serčne & Umali, 1972 | | |-- Cosmonotus White, 1848 | | |-- Notopus De Haan, 1841 | | |-- Ranilia H. Milne Edwards, 1837 | | `-- Umalia Guinot, 1993 | |-- Ranina Lamarck, 1801 [Ranininae de Haan, 1839] | `--o Raninoidinae Lörenthey & Beurlen, 1929 | |-- Notopoides Henderson, 1888 | |-- Notosceles Bourne, 1922 | `-- Raninoides H. Milne Edwards, 1837 `--+--o Cyclodorippoidea Ortmann, 1892 [Tymoloidea Alcock] | |--o Cymonomidae Bouvier, 1897 | | |-- Curupironomus Tavares, 1993 | | |-- Cymonomoides Tavares, 1993 | | |-- Cymonomus A. Milne-Edwards, 1880 | | |-- Cymopolus A. Milne-Edwards, 1880 | | `-- Elassopodus Tavares, 1993 | |?-o Phyllotymolinidae Tavares, 1998 | | |-- Genkaia Miyake & Takeda, 1970 | | |-- Lonchodactylus Tavares & Lemaitre, 1996 | | `-- Phyllotymolinum Tavares, 1993 | `--o Cyclodorippidae Ortmann, 1892 [Tymolidae Alcock] | |--o Cyclodorippinae Ortmann, 1892 | | |-- Clythrocerus A. Milne-Edwards & Bouvier, 1899 | | |-- Corycodus A. Milne-Edwards, 1880 | | |-- Cyclodorippe A. Milne-Edwards, 1880 | | |-- Deilocerus Tavares, 1993 | | |-- Neocorycodus Tavares, 1993 | | |-- Simodorippe Chace, 1940 | | `-- Tymolus Stimpson, 1858 | `--o Xeinostomatinae Tavares, 1992 | |-- Ketamia Tavares, 1992 | |-- Krangalangia Tavares, 1992 | `-- Xeinostoma Stebbing, 1920 `--o Eubrachyura de Saint Laurent, 1980 |?- †Tepexicercinus tlayuaensis Feldmann, Vega et al., 1998 [Tepexicarcinidae Luque, 2015] `==o Heterotremata Guinot, 1977 [Oxystomata, Oxyrhyncha, Cancridea and Brachyrhyncha in partim] |-- Majoidae Samouelle, 1819 `--+--+-- Hymenosomatoidea Macleay, 1838 | `-- Dorippoidea MacLeay, 1838 `--+-- Calappidae Milne Edwards, 1837 `--+--+-- Parthenopidae MacLeay, 1838 | `-- Portunoidea Rafinesque, 1815 `--+-- Trapeziidae Miers, 1886 `--+-- Eriphiidae MacLeay, 1838 |-- Xanthidae MacLeay, 1838 `--+--+-- Aethridae Dana, 1851 | |-- Carpiliidae Ortmann, 1893 | `--+-- Dairidae Ng & Rodriguez, 1986 | `-- Hepatidae Stimpson, 1871 `--+-- Leucosiidae Samouelle, 1819 `--o Thoracotremata Guinot, 1977 [Brachyrhyncha in partim] |-- Ocypodoidea Rafinesque, 1815 (simpukkataskuravut) `-- Grapsoidea MacLeay, 1838
After Martin & Davis, 2001<==o Brachyura (Crabs; taskuravut) |--o Dromiacea | |--o Homolodromioidea | | `-- Homolodromiidae | |--o Dromioidea | | |-- Dromiidae | | `-- Dynomenidae | `--o Homoloidea | |-- Homolidae | |-- Latreilliidae | `-- Poupiniidae `--o Eubrachyura |--o Raninoida | |--o Raninoidea | | |-- Symethidae | | `-- Raninidae | `--o Cyclodorippoidea | |-- Cyclodorippidae | |-- Cymonomidae | `-- Phyllotymolinidae `--+== Heterotremata [paraphyletic] `--o Thoracotremata |--o Pinnotheroidea | `-- Pinnotheriidae |--o Ocypodoidea | |-- Camptandriidae | |-- Mictyridae | |-- Ocypodidae | `-- Palicidae `--o Grapsoidea |-- Gecarcinidae |-- Glyptograpsidae |-- Grapsidae |-- Plagusiidae |-- Sesarmidae `-- Varunidae |
After Parker, 1982, with some families from Martin & Davis, 2001<==o Brachyura |--o Dromiacea | |--o Dromioidea | | |-- Homolodromiidae | | |-- Dromiidae | | |-- Dynomenidae | | |-- Cymonomidae | | `-- Tymolidae [not recognized by Martin & Davis, 2001] | |--o Homoloidea | | |-- Homolidae | | |-- Latreilliidae | | `-- Poupiniidae | `--o Raninoidea | `-- Raninidae |--o Oxystomata | |--o Dorippoidea | | |?- Orithyiidae [from Martin & Davis, 2001] | | `-- Dorippidae | `--o Leucosioidea | |?- Hepatidae [from Martin & Davis, 2001] | |?- Matutidae [from Martin & Davis, 2001] | |-- Calappidae | `-- Leucosiidae |--o Oxyrhyncha | |--o Majoidea | | |?- Epialtidae [from Martin & Davis, 2001] | | |?- Inachidae [from Martin & Davis, 2001] | | |?- Inachoididae [from Martin & Davis, 2001] | | |?- Mithracidae [from Martin & Davis, 2001] | | |?- Pisidae [from Martin & Davis, 2001] | | |?- Tychidae [from Martin & Davis, 2001] | | `-- Majidae | |--o Hymenosomatoidea | | `-- Hymenosomatidae | |--o Mimilambroidea | | `-- Mimilambridae | `--o Parthenopoidea | |?- Aethridae [from Martin & Davis, 2001] | |?- Dairidae [from Martin & Davis, 2001] | |?- Daldorfiidae [from Martin & Davis, 2001] | `-- Parthenopidae |--o Cancridea | `--o Cancroidea | |-- Atelecyclidae | |-- Cancridae | |?- Cheiragonidae [from Martin & Davis, 2001] | |-- Corystidae | |-- Pirimelidae | `-- Thiidae `--o Brachyrhyncha |?-o Cryptochiroidea | `-- Cryptochiridae [from Martin & Davis, 2001] |--o Portunoidea | |-- Ceryonidae | `-- Portunidae |--o Bythograeoidea | `-- Bythograeidae |--o Xanthoidea | |?- Carpiliidae [from Martin & Davis, 2001] | |?- Eumedonidae [from Martin & Davis, 2001] | |-- Goneplacidae | |-- Hexapodidae | |?- Menippidae [from Martin & Davis, 2001] | |?- Panopeidae [from Martin & Davis, 2001] | |?- Pilumnidae [from Martin & Davis, 2001] | |-- Platyxanthidae | |?- Pseudorhombilidae [from Martin & Davis, 2001] | |?- Trapeziidae [from Martin & Davis, 2001] | `-- Xanthidae |--o Bellioidea | `-- Belliidae |--o Grapsidoidea | |-- Grapsidae | |-- Gecarcinidae | `-- Mictyridae |--o Pinnotheroidea | `-- Pinnotheridae |--o Potamoidea [incl. Pseudothelphusoidea; Gecarcinucoidea] | |-- Deckeniidae | |-- Gecarcinucidae [Gecarcinucoidea] | |-- Isolapotamidae [not recognized by Martin & Davis, 2001] | |-- Parathelphusidae [Gecarcinucoidea] | |?- Platythelphusidae [from Martin & Davis, 2001] | |-- Potamidae | |-- Potamocarcinidae [not recognized by Martin & Davis, 2001] | |-- Potamonautidae | |-- Pseudothelphusidae [Pseudothelphusoidea Ortmann, 1893] | |-- Sinopotamidae [not recognized by Martin & Davis, 2001] | |-- Sundathelphusidae [not recognized by Martin & Davis, 2001] | `-- Trichodactylidae [treated as Portunoid by Martin & Davis, 2001] |--o Ocypodoidea | |?- Camptandriidae [from Martin & Davis, 2001] | |?- Mictyridae [from Martin & Davis, 2001] | |-- Ocypodidae | |-- Retroplumidae [placed in Retroplumoidea by Martin & Davis] | `-- Palicidae `--o Hapalocarcinoidea [= Cryptochiroidea (Martin & Davis, 2001)] `-- Hapalocarcinidae [= Cryptochiridae (Martin & Davis, 2001)] |
Reference(s):
- Ahyong, S. T., Lai, J. C. Y., Sharkey, D., Colgan, D. J. & Ng, P. K. L., 2007: Phylogenetics of the brachyuran crabs (Crustacea: Decapoda): The status of Podotremata based on small subunit nuclear ribosomal RNA.
–Molecular Phylogenetics and Evolution: Vol. 45, #2, pp. 576-586 [doi: http://dx.doi.org/10.1016/j.ympev.2007.03.022] - Lahti, S., Malmström, K. K., Koli, L., Leikola, A., Syrjämäki, J. & Lahti, J., 1980: Zoo, Suuri Eläinkirja 6: Selkärangattomat.
–Werner Söderström Osakeyhtiö, Porvoo-Helsinki-Juva, 1980.
Original work [in French]: Beaut, Du Monde Animal XI & XII Invertébrés.
–Rizzoli Editore, Milano, 1968. - Luque, J., 2015: The oldest higher true crabs (Crustacea: Decapoda: Brachyura): insights from the Early Cretaceous of the Americas.
–Palaeontology: Vol. 58, #2, pp. 251-263 [doi: 10.1111/pala.12135] - Martin, J. W. & Davis, G. E., 2001: An Updated Classification of the Recent Crustacea.
–Natural History Museum of Los Angeles County, Science Series: Vol. #39, pp. vii-124 - Parker, S. P. (ed.), 1982: Synopsis and classification of living organisms. Vols. 1 & 2
–McGrew-Hill Book Company