After Siveter, Briggs, Siveter, Sutton, Legg. & Joomun, 2014
<==o Aiolopoda Hou & Bergström, 1996 [incl. Arthropoda von Siebold, 1848][Panarthropoda] (water bears, velvet worms, anomalocarids and arthropods; karhukaiset, käsnäjalkaiset, anomalocaridit ja niveljalkaiset) |?- †Facivermis yunnanicus (transitional form between pripulid/palaeoscolecid worms and lobopods??) |?- †Mongolitubulus squamifer Missarzhevsky, 1977 [sclerites] |-- Tardigrada Spallanzani, 1777 (water bears; karhukaiset) [considered as Aschelmithes by Hou & Bergström (1997)] `==o Loboboda Snodgrass, 1938 [Onychophora sensu lato] [Microdictyoniida; Protonychophora Hutchinson, 1930] (lobopods, velvet worms; käsnäjalkaiset) [paraphyletic grade of earliest arthropods] |-- †Diania cactiformis Liu, Steiner, Dunlop, Keupp, Shu, Ou, Han, Zhang & Zhang, 2011 `--+-- †Paucipodia inermis Chen, Zhou & Ramsköld, 1995 `--+--o †Eoconchariidae Hou & Shu, 1987 [Eoconchariidae Shu & Chen, 1988; Microdictyonidae Chen, Hou & Lu, 1989] | |?- †Fusuconcharium typicum Hao & Shu, 1987 [incl. Fusuconcharium minimum Hao & Shu, 1987] [sclerites] | |?-o †Quadratapora Hao & Shu, 1987 [sclerites] | | |-- †Q. zhenbaensis Hao & Shu, 1987 | | `-- †Q. tenuiporatum (Bengston, Matthews & Missarzhevsky, 1986) [Microdictyon tenuiporatum Bengston, Matthews & Missarzhevsky, 1986] | `--o †Microdictyon Bengston, Matthews & Missarzhevsky in Missarzhevsky & Mambetov, 1981 [Eoconcharium] | |-- †M. chinense (Hao & Shu, 1987) [Eoconcharium chinense Hao & Shu, 1987; Eoconcharium ovalum Hao & Shu, 1987; Eoconcharium spiniferum Hao & Shu, 1987; Eoconcharium rhomboidale Hao & Shu, 1987; Eoconcharium chinense Shu & Shen, 1988; Microdictyon anus Tong, 1989; Microdictyon chinense (Hao & Shu); Zhang & Aldridge, 2007] | |-- †M. effusum Bengston, Matthews & Missarzhevsky in Missarzhevsky & Mambetov, 1981 | |?- †M. rhomboidale Bengston, Matthews & Missarzhevsky, 1986 [Eoconcharium rhomboidale Hao & Shu, 1987] | |-- †M. robisoni | |-- †M. sinicum (Chen, Hou & Lu, 1989) | |-- †M. sphaeroides | |?- †M. depressum Bengston, 1990 | `-- †M. jinshaense Zhang & Aldridge, 2007 `--+-- †Cardiodictyon catenulum Hou, Ramsköld & Bergström, 1991 [†Cardiodyctyidae Hou & Bergström, 1995] `--+--o †Hallucigenia Conway Morris, 1977 [†Hallucigena; †Hallucinogenia] [Hallucigeniidae Conway Morris, 1977] | |-- †H. sparsa Conway Morris, 1977 [M.Camb.] | |-- †H. fortis Hou & Bergström, 1995 [E.Camb.] | `-- †H. hongmeia Steiner, Hu, Liu & Keupp, 2012 [M.Camb.] `--+--o Udeonychophora Poinar, 2000 [Onychophora sensu lato] | `==o “Ontonychophora” Poinar, 2000 [paraphyletic] | |?- †Orstenotubulus evamuellerae Maas, Mayer, Kristensen & Waloszek, 2007 | |-- †Antennacanthopodia gracilis Ou, Liu, Shu, Han, Zhang, Wan & Lei, 2011 | `--+--o †Helenodoridae Poinar, 2000 | | |-- †Helenodorus inopinata Thompson & Jones, 1980 [Carb. NAm.] | | `--o †Ilyodes Scudder, 1890 [Carb. NAm.] | | |-- †I. divisa Scudder, 1890 | | `?- †I. (Helenodorus) opinata Thompson & Jones 1980 [=? Helenodorus inopinata Thompson & Jones, 1980] | `--+-- †Succinipatopsis balticus Poinar, 2000 [L.Eoc. Eu.] [Succinipatopsidae Poinar, 2000] | |-- †Tertiapatus dominicanus Poinar, 2000 [Tert. Caribb.] [Tertiapatidae Poinar, 2000] | `--o Onychophora Grube, 1853 [Euonychophora Hutchinson, 1930 sensu Poinar, 2000] (modern lobopods, velvet worms; nykykäsnäjalkaiset) [10 Recent genera?, 110 species? (90-140)] | |-- Peripatidae Evans, 1902 (South American velvet worms; eteläamerikan käsnäjalkaiset) | `-- Peripatopsidae Bouvier, 1907 (Australian and African velvet worms; Australian ja Afrikan käsnäjalkaiset) `--+--+-- †Onychodictyon ferox Hou, Ramsköld & Bergström, 1991 [†Paronychophora Hou & Bergström, 1995: †Onychodyctyidae Hou & Bergström, 1995] | `--o †Luolishaniidae Hou & Bergström, 1995 [Archonychophora Hou & Bergström, 1995:] | |-- †Acinocricus stichus Conway Morris & Robison, 1988 | |-- “Collins' monster” [Emu Bay Shale] | |-- “Collins' monster” [Burgess Shale] | |-- †Collinsium ciliosum Yang, Ortega-Hernández, Gerber, Butterfield, Hou, Lan & Zhang, 2015 | `-- †Luolishania longicruris Hou & Chen, 1989 `--+-- †Ayshenia pedunculata Walcott, 1911 [Aysheaiidae Walcott 1911] [paraphyletic †Protonychophora Hutchinson, 1930] `==o †Xenusiida Dzik & Krumbiegel, 1989 [paraphyletic †Protonychophora Hutchinson, 1930] |-- †Siberion lenaicus Dzik, 2011 [Siberiida Dzik, 2011: Siberiidae Dzik, 2011] `--+--o †Xenusiidae Dzik & Krumbiegel, 1989 | |-- †Xenusion auerswaldae Pompeckj, 1927 | `-- †Hadranax augustus Budd & Peel, 1998 `--+--+-- †Jianshanopodia decora Liu, Shu, Han, Zhang & Zhang, 2006 | `--o †Megadictyon Luo & Hu, 1999 | |-- †M. haikouensis Luo & Hu, 1999 | `-- †M. cf. haikouensis Liu & Shu, 2007 `--+?- †Miraluolishania haikouensis Liu & Shu, 2004 |-- †Kerygmachela kierkegaardi Budd, 1993 `--+-- †Pambdelurion whittingtoni Budd, 1997 `--o Dinocarida Collins, 1996 sensu Vinther, Stein, Longrich & Harper, 2014 |-- †Opabinia regalis Walcott, 1912 [Opabinida] `--+-- †Caryosyntrips serratus Daley & Budd, 2010; M.Camb. WNAm. |--o †Radiodonta Collins, 1996 sensu Vinther, Stein, Longrich & Harper, 2014 | `-- †Anomalocarida Vinther, Stein, Longrich & Harper, 2014 (anomalocarids; anomalocaridit) `-- Euarthropoda [Arthropoda sensu stricto] [Schizoramia sensu Hou & Bergström, 1997]
After Yang, Ortega-Hernández, Gerber, Butterfield, Hou, Lan & Zhang, 2015<==o Pan-Arthropoda |--o Onychophora sensu lato | |-- †Xenusion | |-- †Diania | `--+-- †Onychodictyon gracilis | `--+-- †Paucipodia | `--+-- †Microdictyon | `--+-- †Cardiodictyon | `--+--+-- †Hallucigenia hongmeia | | `--o †Luolishaniidae | | |-- †Acinocricus | | |-- “Collins' monster” [Emu Bay Shale] | | |-- “Collins' monster” [Burgess Shale] | | |-- †Collinsium | | `-- †Luolishania | `--+-- †Hallucigenia fortis | |-- †Hallucigenia sparsa | `--+-- †Orstenotubulus | `--+-- †Antennacanthopodia | `--o Onychophora sensu stricto | |-- †Tertiapatus | |-- †Ilyodes | |-- Plicatoperipatus | |-- Ooperipatellus | `-- Euperipatoides `--o Tactopoda |--o Tardigrada sensu lato | |-- †Onychodictyon ferox | `-- Tardigrada `--o Euarthropoda sensu lato |--+-- †Jianshanopodia | `-- †Megadictyon `--+-- †Hadranax `--+-- †Kerygmachela `--o †Radiodonta |-- †Pambdelurion `--+-- †Anomalocarida `--o Arthropoda |-- †Opadinia `--o Deuteropoda `-- Euarthropoda sensu stricto? |
After Clowes, 2002– ["Chris’s No-Frills Home Page"], Dzik, 2003, Zhang & Aldridge, 2007, and Liu, Shu, Han, Z. Zhang & X. Zhang, 2008<==o Lobopodia `==o †Microdictyoniida |?- †Facivermis |--o †Archonychophora | `--o †Luolishaniidae | `-- †Luolishania |--o †Paronychophora | `--o †Onychodyctyidae | `-- †Onychodictyon |-- †Paucipodia |--+--o †Scheronychophora | | |--o †Cardiodyctyidae | | | `-- †Cardiodictyon | | |--o †Hallucigeniidae | | | `-- †Hallucigenia | | `-- †Eoconchariidae | | |-- †Fusuconcharium | | |-- †Quadratapora | | `-- †Microdictyon | `--+--o †Xenusiida | | `==o †Protonychophora | | `--o †Xenusiidae | | |-- †Hadranax | | |-- †Jianshanopodia | | `-- †Xenusion | |?- †Mongolitubulus | `-- Tardigrada `--+--+?- †Kerygmachela | |?- †Pambdelurion | |?- †Miraluolishania | |?- †Megadictyon | `--+-- †Radiodonta | `-- Euarthropoda `==o †Protonychophora |--o †Aysheaiidae | `-- †Ayshenia `--o Udeonychophora `==o †Ontonychophora |--o †Helenodoridae | `-- †Helenodorus |--o †Tertiapatidae | `-- †Tertiapatus |--o †Succinipatopsidae | `-- †Succinipatopsis `--o Onychophora |-- Peripatidae `-- Peripatopsidae |
Proposed classification of Poinar, 2000<==o Lobopodia |-- Tardigrada `--o Onychophora |--o †Xenusia | |--o †Archonychophora | | `-- †Luolishaniidae | |--o †Protonychophora | | |-- †Aysheaiidae | | `-- †Xenusiidae | |--o †Scheronychophora | | |-- †Eoconchariidae | | |-- †Hallucigeniidae | | `-- †Cardiodyctyidae | `--o †Paronychophora | `-- †Onychodyctyidae `--o Udeonychophora |--o †Ontonychophora | |--o †Helenodoroidea | | `-- †Helenodoridae | `--o †Tertiapatoidea | |-- †Tertiapatidae | `-- †Succinipatopsidae `--o Onychophora |-- Peripatidae `-- Peripatopsidae |
Reference(s):
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–iNet: Systema Naturae 2000: http://sn2000.taxonomy.nl/Main/Classification/15765.htm - Clowes, C., 2002: Arthropoda
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