After McKenna & Bell, 1997, de Muizon, Cifelli & Céspedes, 1997, and Szalay & Trofimov, 1996, with species from Alroy, 2002

Family -level taxonomy of non-australian metatherians.

Phylogeny of Metatheria after Rougier, Davis & Novacek, 2015

Phylogeny of Metatheria after Vullo et al., 2009

Phylogeny of Metatheria after Case, Goin & Woodburne, 2005

Phylogeny of Metatheria after Szaley & Trofimov, 1996

<==o Metatheria Huxley, 1880 [Marsupialia sensu lato] (marsupials and relatives; pussieläimet)
   |-- †Sinodelphys szalayi Luo, Ji, Wible & Yuan, 2003
   |-- †Holoclemensia texana (Slaughter, 1968) [†Clemensia Slaughter, 1968 (non Clemensia Packard, 1864, a butterfly)] L.Cret. NA. [Holoclemensiidae Aplin & Archer, 1987; Protodelphia Archer, 1984]
   `--+--+-- †Pappotherium pattersoni Slaughter, 1965 sensu Bi, Jin, Li & Du, 2015
      |  `-- †Deltatheroida Kielan-Jaworowska, 1982 [taxonomic history]
      `--o Marsupialiformes Vullo, Gheerbrant, de Muizon & Néraudeau, 2009 [“Neometatheria”]
         |?- †Arcantiodelphys marchandi Vullo, Gheerbrant, de Muizon & Néraudeau, 2009
         |?- †Aenigmadelphys archeri Cifelli & Johanson, 1994, U.Cret. NA.
         |?-o †Anchistodelphys Cifelli, 1990b; U.Cret. NA.
         |  |?- ?†A. delicatus Cifelli, 1990c
         |  `-- †A. archibaldi Cifelli, 1990b
         |?- †Neometatheria genera et species indet. [UA8699] L. Cret. Mad.
         `==o †Boreometatheria (Asiatic and North American metatheres; aasialaiset ja pohjois-amerikkalaiset pussieläimet) [a paraphyletic assemblage of non-Notometatherian metatheres]
            |?- †Camptomus amplus Marsh, 1889
            |?- †Adinodon pattersoni Hershkovitz, 1995 [Adinodontinae Hershkovitz, 1995]
            `--+-- †Iqualadelphis lactea Fox, 1987
               `--+--o †Asiadelphia Trofinov & Szalay, 1994
                  |  |?- †Kokopellia juddi Cifelli, 1993; L.Cret. NA. [Kopellia lapsus calami?]
                  |  `--+-- †Asiatherium reshetovi Trofinov & Szalay, 1994 [Asiatheriidae Trofinov & Szalay, 1994]
                  |     `?-o †Marsasia Nesov, 1997 [?†Asiatherium; jr. synon. Sulestes (Deltatheroida)]
                  |        |-- †M. aenigma Nesov, 1997 
                  |        `-- †M. sp. [Averianov & Kielan-Jaworowska, 1999]
                  `==o ?†Ameridelphia Szalay, in Archer, ed., 1982 sensu stricto? []
                     |?- †Dakotadens morrowi Eaton, 1993b
                     |?- †Progarzonia Ameghino, 1904
                     |?-o †Iugomortiferum Cifelli, 1990b; U.Cret. NA.
                     |  |-- †I. sp. [Cifelli, 1990b] [undiscribed]
                     |  `-- †I. thoringtoni Cifelli, 1990b
                     |?-o †Protalphadon Cifelli, 1990
                     |  |-- †P. lulli (Clemens, 1966) Cifelli, 1990 [Alphadon lulli Clemens, 1966]
                     |  `-- †P. foxi Johanson, 1996b
                     |-- †Varalphadon cerber (Fox, 1971) Johanson, 1996 [Protalphadon cerber (Fox, 1971); Alphadon cerber Fox, 1971]
                     |-- †Varalphadon waheapensis (Cifelli, 1990a) Johanson, 1996 [Protalphadon waheapensis Cifelli, 1990a]
                     |-- †Varalphadon crebreforme (Cifelli, 1990b) Johanson, 1996 [Protalphadon crebreforme Cifelli, 1990b]
                     |--o †Stagodontidae Marsh, 1889 sensu McKenna & Bell, 1997 [taxonomic history] [†Anchimetatheria Szalay, 1993, in partim]
                     |  |?- †Boreodon matutinus Lambe, 1902 (nomen dubium, Fox & Naylor, 2006)
                     |  |?-o †Pariadens Cifelli & Eaton, 1987 [treated as stagodonts by Cifelli & Eaton, 1987 and Cifelli, 2004]
                     |  |  |-- †P. kirklandi Cifelli & Eaton, 1987
                     |  |  `-- †P. mckennai Cifelli, 2004
                     |  `-- †Stagodontidae Marsh, 1889 sensu stricto [taxonomic history] [†Anchimetatheria Szalay, 1993, in partim]
                     |--+--+-- †Bistius bondi Clemens & Lillegraven, 1986
                     |  |  `--o †Turgidodon Cifelli, 1990 [Alphadon Simpson, 1927]
                     |  |     |-- †T. praesagus (Russell, 1952) Cifelli, 1990 [Alphadon praesagus Russell, 1952]
                     |  |     |-- †T. rhaister (Clemens, 1966) Cifelli, 1990 [Alphadon rhaister Clemens, 1966]
                     |  |     |-- †T. russelli (Fox, 1979) Cifelli, 1990 [Alphadon russelli Fox, 1979]
                     |  |     |-- †T. parapraesagus (Rigby & Wolberg, 1987) Cifelli, 1990 [Alphadon parapraesagus Rigby & Wolberg, 1987]
                     |  |     |-- †T. petiminis Storer, 1991
                     |  |     |-- †T. lillegraveni Cifelli, 1990
                     |  |     `-- †T. madseni Cifelli, 1990
                     |  `--+-- †Alphadontidae Marshall, Case & Woodburne, 1990 [Alphadontia Archer, 1984; Didelphidae: Alphadontinae Marshall, Case & Woodburne, 1990 sensu McKenna & Bell, 1997]
                     |     `-- †Pediomyidae Simpson, 1927 [†Anchimetatheria Szalay, 1993, in partim]
                     `--+-- †Jaskhadelphys minutus Marshall & de Muizon, 1988 [Jaskhadelphysidae Marshall & de Muizon, 1988 or de Muizon, 1991]
                        |?-+?- †Thylacodon pusillus Matthew & Granger, 1921 [P. pusillus Matthew & Granger, 1921]
                        |  `?- †Simpsonitheria Szalay, in Szalay, Novacek & McKenna, eds., 1993
                        `--+?- †Herpetotheriidae Trouessant, 1879 [Didelphidae: Herpetotheriinae Trouessant, 1879 sensu McKenna & Bell, 1997] [paraphyletic?]
                           `-- Notometatheria (South American and Australian metatheres, incl. Marsupialia; etelä-amerikkalaiset ja australialaiset pussieläimet)

Metatheria (Marsupialia) incertae sedis:

Garatherium mahboubii Crochet, 1984; E.Eoc. NAf.
    †Kasserinotherium tunisiense Crochet, 1989; E.Eoc. NAf.
    †Ghamidtherium dimaiensis Sánches-Villagra et al., 2007; L.Eoc. NEAf.

Alternative phylogenies of Metatheria:

After Rougier, Davis & Novacek, 2015

<==o Metatheria
   |-- †Holoclemensia
   `--+--+-- †Pappotherium
      |  `-- †Deltatheroida
      `--+--+-- †Asiatherium
         |  `--+-- †Aenigmadelphys 1
         |     `--+---+--+-- †Kokopellia
         |        |   |  `-- †Iugomortiferum
         |        |   `--+-- †Turgidodon
         |        |      `--+-- †Albertatherium
         |        |         `-- †Alphadon
         |        `--+?- †Anchistodelphys
         |           `--+--+-- †Pediomys
         |              |  `-- †Glasbius
         |              `--+?- †Aenigmadelphys 2
         |                 `--+-- †Pariadens
         |                    `--+-- †Eodelphis
         |                       `-- †Didelphodon
         `--+-- Gurlin Tsav Skull
            `--+-- †Borhyaenidae
               `--+-- †Mayulestes
                  `--+-- †Jaskhadelphys
                     `--+-- †Pucadelphys
                        `--+-- †Andinodelphys
                           `--+-- Didelphidae
                              `--+-- Microbiotheria
                                 `-- Dasyuromorphia

Reference(s):

  • Rougier, G. W., Davis, B. M. & Novacek, M. J., 2015: A deltatheroidan mammal from the Upper Cretaceous Baynshiree Formation, eastern Mongolia.
    –Cretaceous Research: Vol. 52, pp. 167-177 [doi: 10.1016/j.cretres.2014.09.009]

After Vullo, Gheerbrant, de Muizon & Néraudeau, 2009

<==o Metatheria
   |-- †Sinodelphys
   `--+-- †Deltatheroida
      `--o Marsupialiformes
         |-- †Arcantiodelphys
         |-- †Dacotadens
         `--+-- †Kokopellia
            `--+-- †Asiatherium
               `--+-- Didelphis
                  `--+-- †Iugomortiferum
                     |-- †Sinbadelphys
                     |-- †Adelodelphys
                     `-- †Pariadens

After Case, Goin & Woodburne, 2005

<==o Metatheria
   |?- †Deltatheroida
   |?-o †Asiadelphia
   |  |?- †Kokopellia
   |  `--+-- †Asiatherium
   |     `-- †Marsasia
   `--o Marsupialia
      |--o †“Alphadelphia”
      |  `--o †Peradectia
      |     |-- †Peradectidae
      |     |-- †Pediomyidae
      |     |-- †Stagodontidae
      |     `-- †Caroloameghiniidae
      |--o Ameridelphia
      |  |?- †Andinodelphys
      |  |--o Didelphimorphia
      |  |  `--o Didelphoidea
      |  |     |-- †Pucadelphidae
      |  |     |-- Didelphidae
      |  |     |-- Caluromyidae
      |  |     |-- †Herpetotheriidae
      |  |     |-- †Derorhynchidae
      |  |     `-- †Protodidelphidae
      |  |-- Paucituberculata
      |  `--o †Sparassodonta
      |     `--o †Borhyaenoidea
      |        |-- †Mayulestidae
      |        |-- †Hondadelphidae
      |        |-- †Hathliacynidae
      |        |-- †Borhyaenidae
      |        |-- †Proborhyaenidae
      |        `-- †Thylacosmilidae
      `--o Australidelphia
         |-- Microbiotheria
         |-- Peramelia
         |-- Dasyuromorphia
         |-- Notorycteromorpha
         `-- Diprotodontia

After Szalay & Trofimov, 1996

<==o Metatheria
   |-- †Deltatheroida
   `--+--o †Asiadelphia
      |  |?- †Kokopellia
      |  `--+-- †Marsasia
      |     `-- †Asiatherium
      `==o †Archimetatheria
         |-- †Stagodontidae
         |-- †Pediomyidae
         |--+-- †Sudameridelphia
         |  `--+-- †Simpsonitheria
         |     `-- Paucituberculata
         `--+-- Didelphimorphia
            `--+--+-- Microbiotheria
               |  `-- Dasyuromorphia
               `--o Syndactyli
                  |--+?- Peramelia
                  |  `?- Notorycteromorpha
                  `--+?-o †Yalkaparidontia
                     |  `-- †Yalkaperidon
                     `-- Diprotodontia

Reference(s):