After McKenna & Bell, 1997, de Muizon, Cifelli & Céspedes, 1997, and Szalay & Trofimov, 1996, with species from Alroy, 2002

Family -level taxonomy of non-australian metatherians.

Phylogeny of Metatheria after Szaley & Trofimov, 1996

Phylogeny of Metatheria after Case, Goin & Woodburne, 2005

Phylogeny of Metatheria after Vullo et al., 2009

<==o Metatheria Huxley, 1880 [Marsupialia sensu lato] (marsupials and relatives; pussieläimet)
   |-- †Sinodelphys szalayi Luo, Ji, Wible & Yuan, 2003
   |-- †Holoclemensia texana (Slaughter, 1968) [†Clemensia Slaughter, 1968 (non Clemensia Packard, 1864, a butterfly)] L.Cret. NA. [Holoclemensiidae Aplin & Archer, 1987; Protodelphia Archer, 1984]
   |-- †Deltatheroida Kielan-Jaworowska, 1982 [taxonomic history]
   `--o Marsupialiformes Vullo, Gheerbrant, de Muizon & Néraudeau, 2009 [“Neometatheria”]
      |?- †Arcantiodelphys marchandi Vullo, Gheerbrant, de Muizon & Néraudeau, 2009
      |?- †Aenigmadelphys archeri Cifelli & Johanson, 1994, U.Cret. NA.
      |?-o †Anchistodelphys Cifelli, 1990b; U.Cret. NA.
      |  |?- ?†A. delicatus Cifelli, 1990c
      |  `-- †A. archibaldi Cifelli, 1990b
      |?- †Neometatheria genera et species indet. [UA8699] L. Cret. Mad.
      `==o †Boreometatheria (Asiatic and North American metatheres; aasialaiset ja pohjois-amerikkalaiset pussieläimet) [a paraphyletic assemblage of non-Notometatherian metatheres]
         |?- †Camptomus amplus Marsh, 1889
         |?- †Adinodon pattersoni Hershkovitz, 1995 [Adinodontinae Hershkovitz, 1995]
         `--+-- †Iqualadelphis lactea Fox, 1987
            `--+--o †Asiadelphia Trofinov & Szalay, 1994
               |  |?- †Kokopellia juddi Cifelli, 1993; L.Cret. NA. [Kopellia lapsus calami?]
               |  `--+-- †Asiatherium reshetovi Trofinov & Szalay, 1994 [Asiatheriidae Trofinov & Szalay, 1994]
               |     `--o †Marsasia Nesov, 1997 [?†Asiatherium]
               |        |-- †M. aenigma Nesov, 1997 
               |        `-- †M. sp. [Averianov & Kielan-Jaworowska]
               `==o ?†Ameridelphia Szalay, in Archer, ed., 1982 sensu stricto? []
                  |?- †Dakotadens morrowi Eaton, 1993b
                  |?- †Progarzonia Ameghino, 1904
                  |?-o †Iugomortiferum Cifelli, 1990b; U.Cret. NA.
                  |  |-- †I. sp. [Cifelli, 1990b] [undiscribed]
                  |  `-- †I. thoringtoni Cifelli, 1990b
                  |?-o †Protalphadon Cifelli, 1990
                  |  |-- †P. lulli (Clemens, 1966) Cifelli, 1990 [Alphadon lulli Clemens, 1966]
                  |  `-- †P. foxi Johanson, 1996b
                  |-- †Varalphadon cerber (Fox, 1971) Johanson, 1996 [Protalphadon cerber (Fox, 1971); Alphadon cerber Fox, 1971]
                  |-- †Varalphadon waheapensis (Cifelli, 1990a) Johanson, 1996 [Protalphadon waheapensis Cifelli, 1990a]
                  |-- †Varalphadon crebreforme (Cifelli, 1990b) Johanson, 1996 [Protalphadon crebreforme Cifelli, 1990b]
                  |--o †Stagodontidae Marsh, 1889 sensu McKenna & Bell, 1997 [taxonomic history] [†Anchimetatheria Szalay, 1993, in partim]
                  |  |?- †Boreodon matutinus Lambe, 1902 (nomen dubium, Fox & Naylor, 2006)
                  |  |?-o †Pariadens Cifelli & Eaton, 1987 [treated as stagodonts by Cifelli & Eaton, 1987 and Cifelli, 2004]
                  |  |  |-- †P. kirklandi Cifelli & Eaton, 1987
                  |  |  `-- †P. mckennai Cifelli, 2004
                  |  `-- †Stagodontidae Marsh, 1889 sensu stricto [taxonomic history] [†Anchimetatheria Szalay, 1993, in partim]
                  |--+--+-- †Bistius bondi Clemens & Lillegraven, 1986
                  |  |  `--o †Turgidodon Cifelli, 1990 [Alphadon Simpson, 1927]
                  |  |     |-- †T. praesagus (Russell, 1952) Cifelli, 1990 [Alphadon praesagus Russell, 1952]
                  |  |     |-- †T. rhaister (Clemens, 1966) Cifelli, 1990 [Alphadon rhaister Clemens, 1966]
                  |  |     |-- †T. russelli (Fox, 1979) Cifelli, 1990 [Alphadon russelli Fox, 1979]
                  |  |     |-- †T. parapraesagus (Rigby & Wolberg, 1987) Cifelli, 1990 [Alphadon parapraesagus Rigby & Wolberg, 1987]
                  |  |     |-- †T. petiminis Storer, 1991
                  |  |     |-- †T. lillegraveni Cifelli, 1990
                  |  |     `-- †T. madseni Cifelli, 1990
                  |  `--+-- †Alphadontidae Marshall, Case & Woodburne, 1990 [Alphadontia Archer, 1984; Didelphidae: Alphadontinae Marshall, Case & Woodburne, 1990 sensu McKenna & Bell, 1997]
                  |     `-- †Pediomyidae Simpson, 1927 [†Anchimetatheria Szalay, 1993, in partim]
                  `--+-- †Jaskhadelphys minutus Marshall & de Muizon, 1988 [Jaskhadelphysidae Marshall & de Muizon, 1988 or de Muizon, 1991]
                     |?-+?- †Thylacodon pusillus Matthew & Granger, 1921 [P. pusillus Matthew & Granger, 1921]
                     |  `?- †Simpsonitheria Szalay, in Szalay, Novacek & McKenna, eds., 1993
                     `--+?- †Herpetotheriidae Trouessant, 1879 [Didelphidae: Herpetotheriinae Trouessant, 1879 sensu McKenna & Bell, 1997] [paraphyletic?]
                        `-- Notometatheria (South American and Australian metatheres, incl. Marsupialia; etelä-amerikkalaiset ja australialaiset pussieläimet)

Alternative phylogenies of Metatheria:

After Szalay & Trofimov, 1996

<==o Metatheria
   |-- †Deltatheroida
   `--+--o †Asiadelphia
      |  |?- †Kokopellia
      |  `--+-- †Marsasia
      |     `-- †Asiatherium
      `==o †Archimetatheria
         |-- †Stagodontidae
         |-- †Pediomyidae Simpson, 1927
         |--+-- †Sudameridelphia
         |  `--+-- †Simpsonitheria
         |     `-- Paucituberculata
         `--+-- Didelphimorphia
            `--+--+-- Microbiotheria
               |  `-- Dasyuromorphia
               `--o Syndactyli Gill, 1871
                  |--+?- Peramelia
                  |  `?- Notorycteromorpha
                  `--+?-o †Yalkaparidontia
                     |  `-- †Yalkaperidon
                     `-- Diprotodontia

After Case, Goin & Woodburne, 2005

<==o Metatheria
   |?- †Deltatheroida
   |?-o †Asiadelphia
   |  |?- †Kokopellia
   |  `--+-- †Asiatherium
   |     `-- †Marsasia
   `--o Marsupialia
      |--o †“Alphadelphia”
      |  `--o †Peradectia
      |     |-- †Peradectidae
      |     |-- †Pediomyidae
      |     |-- †Stagodontidae
      |     `-- †Caroloameghiniidae
      |--o Ameridelphia
      |  |?- †Andinodelphys
      |  |--o Didelphimorphia
      |  |  `--o >Didelphoidea
      |  |     |-- †Pucadelphidae
      |  |     |-- Didelphidae
      |  |     |-- Caluromyidae
      |  |     |-- †Herpetotheriidae
      |  |     |-- †Derorhynchidae
      |  |     `-- †Protodidelphidae
      |  |-- Paucituberculata
      |  `--o †Sparassodonta
      |     `--o †Borhyaenoidea
      |        |-- †Mayulestidae
      |        |-- †Hondadelphidae
      |        |-- †Hathliacynidae
      |        |-- †Borhyaenidae Ameghino, 1894
      |        |-- †Proborhyaenidae Ameghino, 1897
      |        `-- †Thylacosmilidae Riggs, 1933
      `--o Australidelphia
         |-- Microbiotheria
         |-- Peramelia
         |-- Dasyuromorphia
         |-- Notorycteromorpha
         `-- Diprotodontia

After Vullo, Gheerbrant, de Muizon & Néraudeau, 2009

<==o Metatheria
   |-- †Sinodelphys
   `--+-- †Deltatheroida
      `--o Marsupialiformes
         |-- †Arcantiodelphys
         |-- †Dacotadens
         `--+-- †Kokopellia
            `--+-- †Asiatherium
               `--+-- Didelphis
                  `--+-- †Iugomortiferum
                     |-- †Sinbadelphys
                     |-- †Adelodelphys
                     `-- †Pariadens

Reference(s):