After McKenna & Bell, 1997 and Peláez-Campomanes, 2001, with modern species from Nowak, 1991

<==o Xerini Murray, 1866 [taxonomic history] (African Ground Squirrels; afrikan maaoravat)
   |-- †Heteroxerus Schaub, in Stehlin & Schaub, 1951; L.Olig.-U.Mioc. Eu.
   |--o Atlantoxerus Forsyth Major, 1893 [Xerus (Atlantoxerus) Forsyth Major, 1893] [incl. Getuloxerus Lavocat, 1961] U.Mioc.-L.Plioc. Eu. M.Mioc.-R. Af. ()? [atleast 10 fossil species (Pelaez-Campomanes, 2001)]
   |  |-- †A. blacki (de Bruijn, 1967) [Getuloxerus blacki de Bruijn, 1967; incl. Atlantoxerus idubedensis Cuenca Bescós, 1988]
   |  |-- †A. cuencae (Aquilar et al., 1995)
   |  |-- †A. adroveri (de Bruijn & Mein, 1968)
   |  |-- †A. tadlae (Lavocat, 1961) [Getuloxerus tadlae Lavocat, 1961]
   |  |-- †A. rhodius (de Bruijn et al., 1970)
   |  `-- A. getulus (Linnaeus, 1758) (mahgreb or Barbary ground-squirrel)
   |-- †Aragoxerus Cuenca Bescós, 1988; L.Mioc. Eu.
   |-- Spermophilopsis leptodactylus Blasius, 1884; R. As. (Long-clawed ground squirrel; kynsiorava, kynsimaaorava)
   `--o Xerus Hemprich & Ehremberg, 1832 [Sciurus (Xerus) Hemprich & Ehremberg, 1832] [incl. Geosciurus Smith, 1834; Euxerus Thomas, 1909] L.Plioc.-R. Af. (savannioravat)
      |-- X. erythropus Geoffroy, 1809 (lännensavanniorava, geoffreynsavannio.)
      |--+-- †X. daamsi Denys, Viriot, †Daams, Peláez-Campomanes, Vignaud, Andossa & Brunet, 2003
      |  `-- X. rutilus Cretzschmar, 1826 (unstriped Ground Squirrel)
      `--+-- X. inauris Zimmermann, 1780 (cape Ground Squirrel)
         `-- X. princeps Thomas, 1929 (kaokoveld Ground Squirrel or Mountain Ground Squirrel)

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