After Nowak, 1991
<==o Spermophilus F. Cuvier, 1825 [taxonomic history] (ground squirrels, gophers, susliks; siiselit) |-- †S. pattersoni |-- †S. matachicensis |-- †S. shotwelli |-- †S. howelli |-- †S. meltoni |-- †S. rexroadensis (Hibbard, 1941) |-- †S. finlayensis |-- †S. mcgheei |-- †S. boothi |-- †S. johnsoni |-- †S. dotti |-- †S. sp. |?- S. canus (snake River ground squirrel) |?- †(Poliocitellus) Hibbard, 1942 |--o (Colobotis) Brandt, 1844 | |-- S. (C.) major (reddish or large suslik) | |-- S. (C.) erythrogenys (redcheeked suslik) | `-- S. (C.) fulvus (suurihammassiiseli) |--o (Urocitellus) Obolensky, 1927 | |-- S. (U.) undulatus (pohjans., siperiansiiseli) | |-- S. (U.) parryii (napasiiseli, parrynsiiseli) | `-- S. (U.) columbianus (kolumbiansiiseli) |--o (Spermophilus) F. Cuvier, 1825 | |?- †S. (S.) cragini (Hibbard, 1941) [Citellus cragini Hibbard, 1941] | |-- S. (S.) townsendii (townsend's ground squirrel) | |-- S. (S.) vigilis ()? | |-- S. (S.) mollis (great Basin ground squirrel) | |-- S. (S.) washingtoni (washington ground squirrel) | |-- S. (S.) armatus (uinta ground squirrel) | |-- S. (S.) beldingi (beldinginsiiseli) | |--+-- S. (S.) richardsonii (rishardsoninsiiseli) | | `-- S. (S.) elegans (wyoming ground squirrel) | |-- S. (S.) brunneus (idahonsiiseli) | |-- S. (S.) dauricus (daurian suslik) | |-- S. (S.) alashanicus (alashan ground squirrel) | |-- S. (S.) relictus (tiensaninsiiseli) | |-- S. (S.) citellus (siiseli, lännensiiseli) | |-- S. (S.) xanthoprymnus (near East suslik) | |-- S. (S.) suslicus (suslikki, idänsiiseli) | |-- S. (S.) musicus (north Caucasus suslik) | `-- S. (S.) pygmaeus (pikkusiiseli) |--o Ictidomys Allen, 1877 | |-- S. (I.) tridecemlineatus (leopardisiiseli) | |-- S. (I.) mexicanus (meksikonsiiseli) | |-- S. (I.) spilosoma (spotted ground squirrel) | `-- S. (I.) perotensis (perote ground squirrel) |-- (Poliocitellus) franklinii (franklin's ground squirrel) |--o (Otospermophilus) Brandt, 1844 | |-- S. (O.) variegatus (rock squirrel) | |-- S. (O.) beecheyi (kaliforniansiiseli) | |-- S. (O.) atricapillus (baja California rock squirrel) | |-- S. (O.) annulatus (ringtailed ground squirrel) | `-- S. (O.) adocetus (tropical ground squirrel) |--o (Xerospermophilus) Merriam, 1892 | |-- S. (X.) mohavensis (mohave ground squirrel) | `-- S. (X.) tereticaudus (round-tailed ground squirrel) `--o (Callospermophilus) Merriam, 1897 |-- S. (C.) lateralis (kultasiiseli) |-- S. (C.) saturatus (cascade mantled ground squirrel) `-- S. (C.) madrensis (sierra Madre mantled ground squirrel)
Reference(s):
- Albright III, L. B., 1996: Insectivorans, rodents, and carnivores of the Toledo Bend Local Fauna: An Arikareean (Earlies Miocene) assemblage from the Texas Coastal Plain.
–Journal of Vertebrate Paleontology: Vol. 16, #3, pp. 458-473 - Goodwin, H. T. & Hayes, F. E., 1994: Morphologically derived ground squirrels from the Borchers Local Fauna, Meade County, Kansas, with a redescription of ?Spermophilus cragini.
–Journal of Vertebrate Paleontology: Vol. 14, #2, pp. 278-291 - Martin, R. A., Honey, J. G., Peláez-Campomanes, P., Goodwin, H. T., Baskin, J. A. & Zakrzewski, R. J., 2002: Blancan lagomorphs and rodents of the Deer Park assemblages, Meade County, Kansas.
–Journal of Paleontology: Vol. 76, #6, pp. 1072-1090 - Nowak, R. M., 1991: Walker's Mammals of the World; Part 1.
–The Johns Hopkins University Press, Baltimore and London, 1991, xlviii-642-lxiii - Nowak, R. M., 1991: Walker's mammals of the World; Part 2.
–The Johns Hopkins University Press, Baltimore and London, 1991, xii-643-1629