After Steppan, 1996, McKenna & Bell, 1997, and Weksler, 2003, with recent species from Nowak, 1991
<==o Oryzomyini Vorontsov, 1959 |?-o †Megalomys Truessart, 1881 [taxonomic history] Pleist.-R. W.Ind., Pleist. NSA. [Santa Lucia & Martinique, extinct since 19th century] | |-- †M. desmarestii | `-- †M. luciae |--o Scolomys Anthony, 1924; R. SA. | |-- S. melanops (ecuadorinpiikkihiiri) | `-- S. ucayalensis |--o Zygodontomys J.A. Allen, 1897; Pleist.-R. SA., R. CA. | |-- Z. brevicauda | |-- Z. reigi | `-- Z. brunneus `--+--+--o Oryzomys albigularis -species group | | `-- Oryzomys albigularis | |--+--o Oryzomys megacephalus -species group | | | `-- Oryzomys megacephalus | | `--o Oryzomys yunganus -species group | | `-- Oryzomys yunganus | |--+--o Handleyomys Voss, Gomez-Laverde & Pacheco, 2002 | | | |-- H. fuscatus (Allen, 1912) [Aepeomys fuscatus Allen, 1912] | | | `-- H. intectus (Thomas, 1921) [Oryzomys intectus] | | `--+--o Oryzomys alfaroi -species group | | | `-- Oryzomys alfaroi | | `--o Oryzomys melanotis -species group | | `-- Oryzomys melanotis | |--+-- Oryzomys talamancae | | `--o Oryzomys nitidus -species group | | |?- O. nitidus | | |-- Oryzomys macconnelli | | `--+-- Oryzomys lamia | | `-- Oryzomys russatus | `--o Oecomys Thomas, 1906 [Oryzomys (Oecomys) Thomas, 1906] R. CA. SA. | |?- O. cleberi | |?- O. flavicans | |?- O. paricola | |?- O. phaeotis | |?- O. rex | |?- O. roberti | |?- O. rutilus | |?- O. speciosus | |?- O. superans | |-- O. bicolor | |-- O. catharinae | `--+-- O. trinitatis | `--+-- O. concolor | `-- O. mamorae `--+--+-- Oligoryzomys Bangs, 1900 [ Oryzomys (Oligoryzomys) Bangs, 1900] Pleist.-R. SA., R. CA. | `--+--o Neacomys | | |?- N. guianae | | |?- N. pictus | | |?- N. tenuipes | | |-- N. spinosus | | `--+-- N. minutus | | `-- N. musseri | `--+-- Oryzomys balneator | `--o Microryzomys Thomas, 1917 [incl. Thallomyscus Thomas, 1926] [Oryzomys (Microryzomys) Thomas, 1917] | |-- M. minutus | `?- M. altissimus `--+-- Oryzomys polius `--+-- Oryzomys angouva |-- Oryzomys subflavus |--+-- †Lundomys molitor Voss & Carleton, 1993; U.Pleist.-R. SA. | `--+--o Pseudoryzomys Hershkovitz, 1959; Pleist.-R. SA. | | |-- P. wavrini | | `-- P. simplex | `--o Holochilus Brandt, 1835 [Mus (Holochilus) Brandt, 1835; Holochyse Lesson, 1842] M.Pleist.-R. SA. | |-- †H. primigenus Steppan, 1996 | `--+-- H. brasiliensis | |-- H. sciureus[ssp. => H. brasiliensis??] | |-- H. chacarius[ssp. => H. brasiliensis??] | `-- H. magnus `--+--+-- Amphinectomys savamis Malygin, Aniskin, Isaev & Milishnikov, 1994; R. SA. | `--o Nectomys Peters, 1861 [incl. Cercomys F.Cuvier, 1829 (a chimera)] Pleist.-R. SA. [sgen.=> Oryzomys?] | |-- N. squamipes | |?- N. parvipes (eteläamerikanvesirotta) | `?- ?N. palmipes |--o Oryzomys palustris -species group [Oryzomys sensu stricto (Weksler, 2003)] | |-- Oryzomys couesi | `--o Oryzomys palustris (suoriisirotta) | |-- †O. p. antillarum Thomas, 1898 [O. antillarum] | `-- O. p. palustris `--+-- Oryzomys xantheolus `--+--o Nesoryzomys Heller, 1904; R. W.Pac. | |-- N. darwini | |-- N. fernandinae | |-- N. indefessus | `-- N. swarthi `--+-- Sigmodontomys alfari J.A. Allen, 1897; R. CA. SA. [Oryzomys (Sigmodontomys) J.A. Allen, 1897] `--o Melanomys Thomas, 1902; R. CA. [Oryzomys (Melanomys) Thomas, 1902] |-- M. caliginosus |?- M. robustulus `?- M. zunigae
Oryzomyini not treated by Weksler, 2003
Microakodontomys transitorius Hershkovitz 1993; R. SA.
Oryzomys Baird, 1857 [Hesperomys (Oryzomys) Baird, 1857] [incl. Micronectomys Hershkovitz, 1948 (nomen nudum); Macruroryzomys Hershkovitz, 1948 (nomen nudum)] ?U.Mioc.-R. NA. Pleist.-R. SA., R. CA. W.Ind., R. Pac. (Rice rats; riisirotat)
O. aphrastus
O. argentatus
O. auriventer
O. bolivarae
O. buccinatus
O. capito
O. caudatus
O. chapmani
O. devius
O. dimidiatus
O. fulgens
O. galapagoensis
†O. g. galapagoensis
O. g. bauri
O. gorgasi
O. hammondi
O. intermedius
O. keaysi
O. kelloggi
O. legatus
O. levipes
O. melanostoma
O. nelsoni
O. oniscus
†O. pliocaenicus
O. ratticeps
O. rhabdops
O. rostratus
O. saturatior
Reference(s):
- Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698. - Carroll, R. L., 1988: Appendix. 594-648.
in Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698. - McKenna, M. C. & Bell, S. K., (eds.) 1997: Classification of mammals – above the species level.
–Columbia University Press, New York, 1997, xii-631 - Nowak, R. M., 1991: Walker's Mammals of the World; Part 1.
–The Johns Hopkins University Press, Baltimore and London, 1991, xlviii-642-lxiii - Nowak, R. M., 1991: Walker's mammals of the World; Part 2.
–The Johns Hopkins University Press, Baltimore and London, 1991, xii-643-1629 - Steppan, S. J., 1996: A new species of Holochilus (Rodentia: Sigmodontinae) from the Middle Pleistocene of Bolivia and its phylogenetic significance.
–Journal of Vertebrate Paleontology: Vol. 16, #3, pp. 522-530 - Voss, R. S., Gomez-Laverde, M. & Pacheco, V., 2002: A new genus for Aepeomys fuscatus Allen, 1912, and Oryzomys intectus Thomas, 1921: Enigmatic murid rodents from Andean cloud forests.
–American Museum Novitates: #3373, pp. 1-42 - Weksler, M., 2003: Phylogeny of Neotropical oryzomyine rodents (Muridae: Sigmodontinae) based on the nuclear IRBP exon.
-Molecular Phylogenetics and Evolution: Vol. 29, pp. 331-349