After McKenna & Bell, 1997, and Fleagle & Tejedor, 2002

<==o Homunculini Ameghino sensu Fleagle & Tejedor, 2002 [Callicebini Thomas, 1903 sensu McKenna & Bell, 1997] (titit)
   |-- †Homunculus patagonicus Ameghino, 1894
   |--o †Carlocebus Fleagle, 1990
   |  |-- †C. carmenensis Fleagle, 1990
   |  `-- †C. intermedius Fleagle, 1990
   |?- †Paralouatta veronai Rivero & Arredondo, 1991
   `--o Callicebus Thomas, 1903 (titit; titi's)
      |?- C. moloch (tummatiti)
      |?- C. bernardi
      |?- C. brunneus (brown Titi)
      |?- C. caligatus (booted Titi)
      |?- C. cinerascens (ashygrey Titi)
      |?- C. cupreus (coppery Titi)
      |?- C. donacophilus (d'Orbigny's Titi)
      |?- C. dubius (hershkovitz's Titi)
      |?- C. hoffmannsi (hoffmanns' Titi)
      |?- C. modestus (bolivian Titi)
      |?- C. oenanthe (peruvian Mountain Titi)
      |?- C. olallae (olalla's Titi)
      |?- C. stephennasi
      |-- C. torquatus (kaulustiti)
      `--o C. personatus (naamiotiti)
         |-- C. p. personatus ()
         |-- C. p. brunello ()
         |-- C. p. melanochir ()
         `-- C. p. nigrifrons ()

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