After McKenna & Bell, 1997, and Fleagle & Tejedor, 2002
<==o Homunculini Ameghino sensu Fleagle & Tejedor, 2002 [Callicebini Thomas, 1903 sensu McKenna & Bell, 1997] (titit) |?- †Xenothrix mcgregori (Anthony, 1919) Williams & Koopman, 1952 [Xenotrichinae] (Jamaican monkey) |-- †Homunculus patagonicus Ameghino, 1894 |--o †Carlocebus Fleagle, 1990 | |-- †C. carmenensis Fleagle, 1990 | `-- †C. intermedius Fleagle, 1990 |?- †Paralouatta veronai Rivero & Arredondo, 1991 `--o Callicebus Thomas, 1903 (titit; titi's) |?- C. moloch (tummatiti) |?- C. bernardi |?- C. brunneus (brown Titi) |?- C. caligatus (booted Titi) |?- C. cinerascens (ashygrey Titi) |?- C. cupreus (coppery Titi) |?- C. donacophilus (d'Orbigny's Titi) |?- C. dubius (hershkovitz's Titi) |?- C. hoffmannsi (hoffmanns' Titi) |?- C. modestus (bolivian Titi) |?- C. oenanthe (peruvian Mountain Titi) |?- C. olallae (olalla's Titi) |?- C. stephennasi |-- C. torquatus (kaulustiti) `--o C. personatus (naamiotiti) |-- C. p. personatus () |-- C. p. brunello () |-- C. p. melanochir () `-- C. p. nigrifrons ()
Reference(s):
- Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698 - Fleagle, J. G. & Tejedor, M. F., 2002: Early platyrrhines of southern South America. 161-173.
in Hartwig, W. C., (eds.) 2002: The Primate Fossil Record.
–Cambridge University Press, Cambridge, England, 2002, xiv-530 - Carroll, R. L., 1988: Appendix. 594-648
in Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698 - McKenna, M. C. & Bell, S. K., (eds.) 1997: Classification of mammals – above the species level.
–Columbia University Press, New York, 1997, xii-631 - Nowak, R. M., 1991: Walker's Mammals of the World; Part 1.
–The Johns Hopkins University Press, Baltimore and London, 1991, xlviii-642-lxiii - Nowak, R. M., 1991: Walker's mammals of the World; Part 2.
–The Johns Hopkins University Press, Baltimore and London, 1991, xii-643-1629