Mikko's Phylogeny Archive Mammalian Groups   Rhinocerotidae - rhinos

After Heissig, 1989, McKenna & Bell, 1997 and Alroy, 2002

<==o Rhinocerotidae Owen, 1845 (sarvikuonot)
   |?- †Symphyssorrhachis
   |?- †Meschotherium
   |?- †Punjabitherium
   |?-+-- †Uintaceras radinskyi [=? Teletaceras radinskyi]
   |  `--o †Teletaceras Hanson, 1989 [Eotrigonias Stock, 1949]
   |     |-- †T. radinskyi Hanson, 1989
   |     |-- †T. mortivallis (Stock, 1949)
   |     `-- †T. borissiaki (Beliaeva, 1959)
   |?- †Epiaceratherium bolcense
   `--+--+?-o †Penetrigonias
      |  |  |-- †P. dakotensis (Peterson, 1920) Prothero, 1998b
      |  |  |-- †P. hudsoni
      |  |  `-- †P. sagittatus
      |  `--+--+-- †Amphicaenopus platycephalus (Osborn & Wortman, 1894) Wood, 1927
      |     |  |--o †Ronzotherium Aymard, 1854
      |     |  |  |-- †R. velaunum
      |     |  |  |-- †R. filholi (Osborn, 1900)
      |     |  |  |-- †R. romani
      |     |  |  `-- †R. brevirostre
      |     |  `--o †Trigonias Lucas, 1900
      |     |     |-- †T. osborni Lucas, 1900
      |     |     |-- †T. wellsi
      |     |     `-- †T. yoderensis
      |     `--+-- †Gulfoceras westfalli
      |        `--o †Diceratheriinae Dollo, 1885 [Elasmotheriinae: Diceratheriini Dollo, 1885 sensu Antoine, 2002]
      |           |--o †Subhyracodon [Caenopus, Leptaceratherium]
      |           |  |-- †S. mitis
      |           |  |-- †S. occidentalis
      |           |  `-- †S. trigonodus
      |           `--o †Diceratherium
      |              |-- †D. armatum
      |              |-- †D. annectens
      |              `-- †D. niobrarense
      `--+--o †Aceratheriinae
         |  |-- †Pleuroceros pleuroceros [paraphyletic Menoceratini Prothero, Manning & Hanson, 1986]
         |  `--+-- †Protaceratherium albigense [paraphyletic Menoceratini Prothero, Manning & Hanson, 1986]
         |     `--+-- †Protaceratherium? minutum (Cuvier, 1822) Abel, 1910 [paraphyletic Menoceratini Prothero, Manning & Hanson, 1986]
         |        `--+--o †Menoceras Troxell, 1921 [Moschoedestes] [paraphyletic Menoceratini Prothero, Manning & Hanson, 1986]
         |           |  |-- †M. arikarensis
         |           |  `-- †M. barbouri
         |           `--+-- †Aceratheriini Dollo, 1885 [Aceratheriinae Dollo, 1885 sensu stricto ( McKenna & Bell, 1997)]
         |              `-- †Teleoceratini [Rhinocerotinae: Teleoceratini sensu McKenna & Bell, 1997]
         `--o Rhinocerotinae Owen, 1845 [Rhinocerotini Owen, 1845 sensu McKenna & Bell, 1997] (nykysarvikuonot)
            |-- Dicerorhinini [Dicerorhinina sensu McKenna & Bell, 1997] (villasarvikuonot; woolly rhinos)
            `--+-- †Elasmotheriini Bonaparte, 1845  [Elasmotheriinasensu McKenna & Bell, 1997] (elasmoteerit, elasmotheres)
               `--+-- Rhinocerotini Owen, 1845 [Rhinoceratina Owen, 1845 sensu McKenna & Bell, 1997] (aasiansarvikuonot; asian rhinos)
                  `-- Dicerotini [Dicerotina sensu McKenna & Bell, 1997] (afrikansarvikuonot; african rhinos)

Reference(s):

Alroy, J., 2002: North American fossil mammal systematics database.
–iNet: North American fossil mammal systematics database: http://www.nceas.ucsb.edu/~alroy/nafmsd.html

Hanson, C. B., 1989: Teletaceras radinskyi, a new primitive rhinocerotid from the Late Eocene Clarno Formation of Oregon. 379-398.
in Prothero, D. R. & Schoch, R. M., (eds.) 1989: The Evolution of Perissodactyls.
–Oxford University Press, New York, New York & Oxford, England, 1989, ix-537

Heissig, K., 1989: The Rhinocerotidae. 399-417.
in Prothero, D. R. & Schoch, R. M., (eds.) 1989: The Evolution of Perissodactyls.
–Oxford University Press, New York, New York & Oxford, England, 1989, ix-537

Froelich, D. J., 1999: Phylogenetic systematics of basal perissodactyls.
–Journal of Vertebrate Paleontology: Vol. 19, #1, pp. 140 - 159

McKenna, M. C. & Bell, S. K., (eds.) 1997: Classification of mammals – above the species level.
–Columbia University Press, New York, 1997, xii-631

Prothero, D. R. & Schoch, R. M. 1989: Classification of the Perissodactyla. 530-537.
in Prothero, D. R. & Schoch, R. M., (eds.) 1989: The Evolution of Perissodactyls.
–Oxford University Press, New York, New York & Oxford, England, 1989, ix-537