After McKenna & Bell, 1997, Badiola, Astibia, Pereda Suberbiola & Murelaga, 2002
<==o †Palaeotheriidae Bonaparte, 1850 [Palaeotheroidea, Pachynolophoidea, Pachynolophidae in partim] |?- †Bepitherium jordifusalbae Checa & Colombo, 2004 |?- †Lophiotherium Gervais, 1849 |?- †Paranchilophus Casanovas & Santafé, 1989 |-- †Hallensia `--+-- †Hyracotherium leporinum Owen, 1841 `--+--o †Propachynolophus Lemoine, 1891 | |-- †P. gaudryi | `-- †P. maldani `--+--o †Pachynolophus Pomel, 1847 [Pachynolophinae Paulow, 1877] | |-- †P. duvali | `-- †P. livinierensis `--o †Palaeotheriinae Bonaparte, 1850 |?- †Pseudopalaeotherium longirostratum Franzen, 1872 |?- †Paraplagiotherium Depéret, 1917 |?-o †Leptolophus Remy, 1965 [Pachynolophidae] | |-- †L. stehlini Remy, 1965 | |-- †L. nouleti (Stehlin, 1904) | `-- †L. magnus Remy, 1998 |?- †Cantabrotherium Casanova-Cladellas & Santafé-Llopis, 1987 |-- †Lophiotherium Gervais, 1849 `--+--o †Propalaeotherium Gervais, 1849 | |-- †“P.” messelense | |-- †P. parvulum | |-- †P. argentonicum | |-- †P. helveticum | |-- †P. isselanum (Cuvier, 1824) | `-- †P. hassiacum `--+--o †Plagiolophus Pomel, 1847 [Paloplotherium] [Pachynolophidae] | |-- †P. mazateronensis Cuesta, 1994 | |-- †P. minor | |-- †P. fraasi | `-- †P. annectens `--+-- †Anchilophus depereti Gervais, 1852 `--o †Palaeotherium Cuvier, 1804 |-- †P. castrense |-- †P. ruetimeyeri |-- †P. magnum |-- †P. crassum |-- †P. medium |-- †P. curtum `?- †P. (Franzenitherium) lautricense
Reference(s):
- Badiola, A., Astibia, H., Pereda Suberbiola, X. & Murelaga, X., 2002: First record of the genus Leptolophus Remy, 1965 (Mammalia, Perissodactyla) in the late Eocene (Priabonian) of Europe.
–Geodiversitas: Vol. 24, #4, pp. 841-848 - Checa, L. & Colombo, F., 2004: A new early Eocene palaeothere (Mammalia, Perissodactyla) from northeastern Spain.
–Journal of Vertebrate Paleontology: Vol. 24, #2, pp. 507-512 - Froehlich, D. J., 1999: Phylogenetic systematics of basal perissodactyls.
–Journal of Vertebrate Paleontology: Vol. 19, #1, pp. 140 - 159 - Hooker, J. J., 1989: Character polarities in early perissodactyls and their significance for Hyracotherium and infraordinal relationships. 79-101.
in Prothero, D. R. & Schoch, R. M., (eds.) 1989: The evolution of perissodactyls.
–Oxford University Press, New York, New York & Oxford, England, 1989, ix-537 - Remy, J. A., 2001: Sur le crâne de Propalaeotherium isselanum (Mammalia, Perissodactyla, Palaeotheriidae) de Pépieux (Minervois, Sud de la France).
[About the skull of Propalaeotherium isselanum (Mammalia, Perissodactyla, Palaeotheriidae) from Pépieux, southern France.]
–Geodiversitas: Vol. 23, #1, pp. 105-127