After Froelich, 1999 [from Maas, Hussain, Leinders & Thewissen, 2001]
<==o †Isectolophidae Peterson, 1915 [1919 (Schoch, 1989, Lucas, Holbrook & Emry, 2003)] |?- †Sastrilophus dehmi Sahni & Khare, 1971; M.-L.?Eoc. Ind. |?-o †Gandheralophus Missiaen & Gingerich, 2012; E.Eoc. Ind. | |-- †G. minor Missiaen & Gingerich, 2012 | `-- †G. robustus Missiaen & Gingerich, 2012 |--o †Orientolophus Ding, 1993; E.Eoc. CAs. | |-- †O. hengdongensis | `-- †O. namadicus (Dashzeveg, 1979b) Hooker & Dashzeveg, 2004? [Homogalax namadicus Dashzeveg, 1979b; ?Orientolophus namadicus (Dashzeveg) Ting, 1993; ?Orientolophus namadicus (Dashzeveg) Ting, 1998, ?Orientolophus namadicus (Dashzeveg) Hooker & Dashzeveg, 2004] `--+--+-- †Cymbalophus cuniculus Hooker, 1984; ?E.Eoc. As., E.Eoc. NAm. ["Hyracotherium" cuniculum] | `-- †Systemodon tapirinum ["Hyracotherium" tapirinum] `--+-- †Karagalax mamikhelensis Maas, Hussain, Leinders & Thewissen, 2001 `--+-- †Kalakotia simplicidentata Ranga Rao, 1972 [Aulaxolophus quadrangularis, Chasmotherium mckennai, Schlosseria radinskyi] `--+--o †Cardiolophus Gingerich, 1991; E.Eoc. NAm. | |-- †C. radinskyi Gingerich, 1991 [Homogalax planeticus Gingerich 1980] | `-- †C. semihians (Cope 1882) `--+--o †Homogalax Hay, 1899; E.-M.Eoc. As., E.Eoc.NAm. | |-- †H. aureus Gingerich 1991 | |-- †H. protapirinus (Wortman, 1896) [Systemodon primaevus Wortman, 1896; Homogalax tapirinus (Wortman, 1896)?] | `-- †H. wutuensis `--+--o †Isectolophus Scott & Osborn, 1887; M.Eoc. NAm. [taxonomic history] | |-- †I. radinskyi | |-- †I. latidens (Osborn, Scott & Spier, 1878) [taxonomic history] | `-- †I. annectens Scott & Osborn, 1887 `-- Rhinotapiromorpha Prothero & Schoch, 1989 (sarvikuono-tapiirimaiset kavioeläimet)
Reference(s):
- Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698. - Carroll, R. L., 1988: Appendix. 594-648.
in Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698. - Froelich, D. J., 1999: Phylogenetic systematics of basal perissodactyls.
–Journal of Vertebrate Paleontology: Vol. 19, #1, pp. 140 - 159 - Gazin, C. L., 1976: Mammalian faunal zones of the Bridger Middle Eocene.
–Smithsonian Contributions to Paleobiology: No. 26, pp. iv-25 - Holbrook, L. T., Lucas, S. G. & Emry, R. J., 2004: Skulls of the Eocene perissodactyls (Mammalia) Homogalax and Isectolophus.
–Journal of Vertebrate Paleontology: Vol. 24, #4, pp. 951-956 - Lucas, S. G., Holbrook, L. T. & Emry, R. J., 2003: Isectolophus (Mammalia, Perissodactyla) from the Eocene of the Zaysan Basin, Kazakstan and its biochronological significance.
–Journal of Vertebrate Paleontology: Vol. 23, #1, pp. 238-243 - Maas, M. C., Hussain, S. T., Leinders, J. J. M. & Thewissen, J. G. M., 2001: A new isectolophid tapiromorph (Perissodactyla, Mammalia) from the Early Eocene of Pakistan.
–Journal of Paleontology: Vol. 75, #2, pp. 407-417 - McKenna, M. C. & Bell, S. K., (eds.) 1997: Classification of mammals – above the species level.
–Columbia University Press, New York, 1997, xii-631 - Missiaen, P. & Gingerich, P. D., 2012: New Early Eocene Tapiromorph Perissodactyls from the Ghazij Formation of Pakistan, with Implications for Mammalian Biochronology in Asia.
–Acta Palaeontologica Polonica: Vol. 57, #1, pp. 21-34 [doi: 10.4202/app.2010.0093] - Prothero, D. R. & Schoch, R. M., 1989: Classification of the Perissodactyla. 530-537.
in Prothero, D. R. & Schoch, R. M., (eds.) 1989: The evolution of perissodactyls.
–Oxford University Press, New York, New York & Oxford, England, 1989, ix-537 - Thewissen, J. G. M., Williams, E. M. & Hussain, S. T., 2001: Eocene mammals faunas from Northern Indo-Pakistan.
–Journal of Vertebrate Paleontology: Vol. 21, #2, pp. 347-366 - Schoch, R. M., 1989: A review of the tapiroids. 298-320.
in Prothero, D. R. & Schoch, R. M., (eds.) 1989: The evolution of perissodactyls.
–Oxford University Press, New York, New York & Oxford, England, 1989, ix-537