After Butler, 1978, Coombs, 1989, Lucas & Schoch, 1989, Prothero & Schoch, 1989, McKenna & Bell, 1997 Coombs, Hunt, Stepleton, Albright & Fremd, 2001, and Hooker & Dashzeveg, 2004]
<==o †Chalicotheriidae Gill, 1872 sensu Hooker & Dashzeveg, 2004 [incl. Eomeropidae Gill, 1872 (non Matthew, 1929?)] [Chalicotheroidea Gill, 1872 sensu Lucas & Schoch, 1989, Prothero & Schoch, 1989, McKenna & Bell, 1997 Coombs, Hunt, Stepleton, Albright & Fremd, 2001]
|?- †Olsenia Matthew & Granger, 1925 [paraphyletic Eomeropidae Gill, 1872 (non Matthew, 1929?)]
|--o †Litolophus Matthew & Granger, 1925 [non Radinsky, 1964?] E.-M.? Eoc. Ind. CAs. [paraphyletic Eomeropidae Gill, 1872 (non Matthew, 1929?)]
| |-- †L. gobiensis Matthew & Granger, 1925
| `-- †L. ghazijensis Missiaen & Gingerich, 2012
`--+?- †Danjiangia pingi Wang, 1995 [has been treated as brontothere (Brontotheroidea)]
|?-o †Eomoropus Osborn, 1913 [paraphyletic Eomeropidae Gill, 1872 (non Matthew, 1929?)]
| |-- †E. amarorum (Cope, 1881) Osborn, 1913 [“Triplopus” amarorum Cope, 1881; E. annectens Peterson, 1919]
| |-- †E. quadridentatus Zdansky, 1930 [E. minimus Zdansky, 1930]
| `-- †E. pawnyunti Remy, Jaeger, Chaimanee, Soe, Marivaux, Sudre, Tun, Marandat & Dewaele, 2005
|?-o †Grangeria Zdansky, 1930 [?Lunania Chow, 1957] [paraphyletic Eomeropidae Gill, 1872 (non Matthew, 1929?)]
| |-- †G. canina Zdansky, 1930 [Eomoropus major Zdansky, 1930]
| `-- †G. anarsius (Gazin, 1959) [Eomoropus anarsius Gazin, 1959]
`--o †Chalicotheriidae Gill, 1872 sensu stricto?
|--o †Chalicotheriinae Gill, 1872
| |?- †“Macrotherium” macedonicum Bonis et al., 1995
| |-- †Kalimantsia bulgarica Geraads, Spassov & Kovachev, 2001
| `--o †Chalicotherium Kaup, 1833 [Macrotherium Lartet, in Blainville, 1837]
| |?- †C. giganteum Pictet, 1844
| |-- †C. rusingense Butler, 1962
| `--+-- †C. pilgrimi
| |?- †C. wetzleri
| `--+--+-- †C. salinum
| | `--o †Anisodon Lartet, 1851 [Chalicotherium]
| | `-- †A. grande (Blainville, 1849) [Chalicotherium (Macrotherium) grande]
| `--+-- †C. brevirostris Colbert, 1934
| `--+-- †C. wuduensis
| |-- †C. goldfussi Kaup, 1833
| `--o †Nestoritherium Kaup, 1859
| |-- †N. sinense
| `-- †N. sivalense
`--o †Schizotheriinae Holland & Peterson, 1913
|--o †Schizotherium Gill, 1872
| |-- †S. priscum [“Chalicotherium” modicum; Limognitherium ingens; “Ankylotherium” gaudryi]
| |-- †S. turgaicum [Kyzylkakhippus orlovi]
| |-- †S. chucuae
| |-- †S. avitum
| |-- †S. nabanensis
| `-- †S. ordosium Hu, 1959
`--+-- †Borissiakia betpakdalensis (Flerov, 1938) Butler, 1965 [Meropus betpakdalensis Flerov, 1938; Phylotillon betpakdalensis (Flerov, 1938) Borissiak, 1946]
`--+--o †Phylotillon Pilgrim, 1910
| |-- †P. naricus
| `-- †P. bavaricum (von Koenigswald, 1932)
`--+-- †Tylocephalonyx skinneri Coomps, 1979
|--o †Moropus Marsh, 1877 [Lophiodon]
| |-- †M. oregonensis (Leidy, 1873) [Moropus distans Marsh, 1877; Lophiodon? oregonensis Leidy, 1873]
| |-- †M. elatus
| |-- †M. hollandi
| |-- †M. matthewi
| `-- †M. merriami
`--o †Ancylotherium Gaudry, 1863 [Metaschizotherium; Postschizotherium]
|?- †“Huanghotherium” anlungense Tung, Huang & Qiu, 1975
|?- †“Gansuodon” pingliangense Wu & Chen, 1976
|?- †“Chemositia” tugenensis Pickford, 1979
|?- †A. cheboitense Guérin & Pickford, 2005
|-- †(Metaschizotherium) fraasi von Koenigswald, 1932
`--o †(Ancylotherium) Gaudry, 1863
|-- †A. hennigi (Dietrich, 1942)
`-- †A. pentelicum (Gaudry & Lartet, 1856)
Reference(s):
- Butler, P. M., 1978: Chalicotheriidae. 368-370.
in Maglio, V. J. & Cooke, H. B. S., (eds.) 1978: Evolution of African mammals.
–Harvard University Press, Cambridge, Massachusetts & London, England, 1978, xiv-641
- Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698.
- Carroll, R. L., 1988: Appendix. 594-648.
in Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698.
- Coombs, M. C., 1989: Interrelationships and diversity in the Chalicotheriidae. 438-457.
in Prothero, D. R. & Schoch, R. M., (eds.) 1989: The Evolution of Perissodactyls.
–Oxford University Press, New York, New York & Oxford, England, 1989, ix-537
- Coombs, M. C., Hunt, Jr, R. M., Stepleton, E., Albright III, L. B. & Fremd, T. J., 2001: Stratigraphy, chronology, biogeography, and taxonomy of Early Miocene
small chalicotheres in North America.
–Journal of Vertebrate Paleontology: Vol. 21, #3, pp. 607-620
- Geraads, D., Spassov, N. & Kovachev, D., 2001: New Chalicotheriidae (Perissodactyla, Mammalia) from the Late Miocene of Bulgaria.
–Journal of Vertebrate Paleontology: Vol. 21, #3, pp. 569-606
- Guérin, C. & Pickford, M., 2005: Ancylotherium cheboitense nov. sp., nouveau Chalicotheriidae (Mammalia, Perissodactyla) du Miocène supérieur des Tugen Hills (Kénya).
[Ancylotherium cheboitense nov. sp., new Chalicotheriidae (Mammalia, Perissodactyla) from the Late Miocene of the Tugen Hills (Kenya).]
–Comptes Rendus de l'Académie des Sciencies, Paris: Palevol: Vol. 4, #3, pp. 225-234 [doi: http://dx.doi.org/10.1016/j.crpv.2004.10.005]
- Hooker, J. J. & Dashzeveg, D., 2004: The origin of chalicotheres (Perissodactyla, Mammalia)
–Palaeontology: Vol. 47, #6, pp. 1363-1386
- Lucas, S. G. & Schoch, R. M., 1989: Taxonomy and biochronology of Eomoropus and Grangeria, Eocene chalicotheres from the western
United States and China. 422-437.
in Prothero, D. R. & Schoch, R. M., (eds.) 1989: The Evolution of Perissodactyls.
–Oxford University Press, New York, New York & Oxford, England, 1989, ix-537
- McKenna, M. C. & Bell, S. K., (eds.) 1997: Classification of mammals – above the species level.
–Columbia University Press, New York, 1997, xii-631
- Missiaen, P. & Gingerich, P. D., 2012: New Early Eocene Tapiromorph Perissodactyls from the Ghazij Formation of Pakistan, with Implications for Mammalian Biochronology in Asia.
–Acta Palaeontologica Polonica: Vol. 57, #1, pp. 21-34 [doi: 10.4202/app.2010.0093]
- Prothero, D. R. & Schoch, R. M., 1989: Classification of the Perissodactyla. 530-537.
in Prothero, D. R. & Schoch, R. M., (eds.) 1989: The Evolution of Perissodactyls.
–Oxford University Press, New York, New York & Oxford, England, 1989, ix-537
- Remy, J.-A., Jaeger, J.-J., Chaimanee, Y., Soe, U. A. N., Marivaux, L., Sudre, J., Tun, S. T., Marandat, B. & Dewaele, E., 2005: A new chalicothere from the Pondaung Formation (late Middle Eocene) of Myanmar.
–Comptes Rendus de l'Académie des Sciencies, Paris: Palevol: Vol. 4, pp. 341-349