After McKenna & Bell, 1997, Reguero, Ubilla & Perea, 2003, and Reguero & Castro, 2004
<==o †Typotheria Zittel, 1893 [Hegetotheria Simpson, 1945] (typotheres; typoteerit) |-- †Acama |-- †Brandmayria |--o †Typotheroidea Zittel, 1893 | `-- †Interatheriidae Ameghino, 1887 `--o †Hegetotheroidea Simpson, 1945 |-- †Campanorco inauguralis |--o †Archaeohyracidae | |-- †Archaeohyrax patagonicus Ameghino, 1897 | |-- †Archaeotypotherium propheticus Ameghino, 1897 | |-- †Bryanpattersonia | |-- †Eohegototherium | |-- †Eohyrax | |-- †Protarchaeohyrax gracilis Roth, 1903 | `-- †Pseudhyrax |--o †Hegetotheriidae Ameghino, 1894 [Protypotheriidae Ameghino] | |?- †Paedsotherium | |-- †Pachyruchinae Kraglievich | `--o †Hegetotheriinae Ameghino, 1894 | |-- †Sallatherium altiplanense Reguero & Cerdeño, 2005; L.Olig. SAm. | |--o †Prohegetotherium Ameghino, 1897 [Propachyrucos Ameghino, 1897 (in partim); ?Propachyrucos Kraglievich, 1932; Ethegotherium Simpson, Minoprio & Patterson, 1962] L.Olig. SAm. | | |-- †P. sculptum Ameghino, 1897 [Propachyrucos crassus Ameghino, 1897; Prohegetotherium shumwayi Loomis, 1914] | | `-- †P. schiaffinoi (Kraglievich, 1932) [Propachyrucos? schiaffinoi Kraglievich, 1932; Prohegetotherium carettai Minoprio, 1947; Ethegotherium carettai (Minoprio, 1947) Simpson, Minoprio & Patterson, 1962] | |--o †Hegetotherium Ameghino [Selatherium] E.Mioc.-M.Mioc. SAm. | | `-- †H. mirabile (Ameghino, 1887a) | `-- †Hemihegetotherium Rovereto [incl. Pseudohegetotherium]; L.Mioc. SAm. `--o †Mesotheriidae Alston, 1876 |?-o †Altitypotherium E.Mioc. SAm. | |-- †A. chucalensis | `-- †A. paucidens |?- †Caraguatypotherium munozi M.-L.Mioc. SAm. |?- †Hypsitherium bolivianum E.Plioc. SAm. |?- †Proedium Ameghino, 1895 [Isoproedrium, Proedrium] |?- †Typothericulus |?- †Fiandraia romeroi [Fiandraiinae] `==o †“Trachytheriinae” (Ameghino, 1894) Simpson, 1945 [‘Trachytheridae’ Ameghino, 1894] [paraphyletic] |-- †“Trachytherus” mendocensis Simpson & Minoprio, 1949 `--+-- †Anatrachytherus soriai Reguero & Cerdeño, 2005 [Eutrachytherus Ameghino, 1897 (nomen illegit. pro Trachytherus); Proedrium Ameghino,1897 [lapsus pro Proedium); Ameghinotherium Podestá, 1899; Coresodon Ameghino, 1901 (in partim); Isoproedrium Ameghino, 1904 (nomen illegit. pro Proedium) `--+--o †Trachytherus Ameghino, 1889 [Eutrachytherus] L.Eoc.-L.Olig. SAm. | |-- †T. alloxus | |-- †T. curuzucuatiensis (Podestá, 1899) [‘Ameghinotherium’ curuzucuatiense Podestá, 1899] | |-- †T. spegazzinianus Ameghino, 1889 | `-- †T. subandinus Villarroel, Sempere & Marshall, 1994 `--+-- †Microtypotherium choquecotense M.-L.Mioc. SAm. `--+--o †Eutypotherium [Tachtypotherium, Typothericulus] E.-M.Mioc. SAm. | |-- †E. chico | |-- †E. lehmannnitschei | |-- †E. roveretoi | `-- †E. superans `--o †Mesotheriinae (Alston, 1876) Simpson, 1945 |--o †Typotheriopsis [?Acrotypotherium] L.Mioc. SAm. | |-- †T. chasicoensis | `-- †T. silveyrai `==o †Pseudotypotherium Villarroel, 1974; L.Mioc.-M.Pleist. SAm. |-- †P. achirense Villarroel, 1974 |?- †P. majus |?- †P. minus Villarroel, 1978 `--+--+-- †P. exiguum | `--+-- †P. hystatum [Mesotherium hystatum] | `-- †P. subinsigne `--o †Mesotherium Serres, 1867; E.-M.Pleist. SAm. |-- †M. pachygnathum `--+-- †M. maendrum `-- †M. cristatum
Reference(s):
- Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698. - Carroll, R. L., 1988: Appendix. 594-648.
in Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698. - Cerdeño, E., Bond, M., 1998: Taxonomic revision and phylogeny of Paedotherium and Tremacyllus (Pachyruchinae, Hegetotheriidae, Notoungulata) from the Late Miocene to the Pleistocene of Argentina.
–Journal of Vertebrate Paleontology: vol. 18, #4, pp. 799-811 - Croft, D. A., 2007: The Middle Miocene (Laventan) Quebrada Honda fauna, southern Bolivia and a description of its notoungulates.
–Palaeontology: Vol. 50, #1, pp. 277-303 [doi: 10.1111/j.1475-4983.2006.00610.x] - McKenna, M. C. & Bell, S. K., (eds.) 1997: Classification of mammals – above the species level.
–Columbia University Press, New York, 1997, xii-631 - Reguero, M. A. & Cerdeño, E., 2005: New Late Oligocene Hegetotheriidae (Mammalia, Notoungulata) from Salla, Bolivia.
–Journal of Vertebrate Paleontology: Vol. 25, #3, pp. 674-684 - Reguero, M. A. & Castro, P. V., 2004: Un nuevo Trachytheriinae (Mammalia,?Notoungulata) del Deseadense (Oligoceno tardío) de Patagonia, Argentina: implicancias en la filogenia, biogeografía y bioestratigrafía de los Mesotheriidae.
[A new Trachytheriinae (Mammalia, †Notoungulata) from the Deseadan (Late Oligocene) of Patagonia, Argentina: phylogeny, biogeography, and biostratigraphy of the Mesotheriidae Family.]
–Revista Geologica de Chile: Vol. 31, pp. 45-64 [doi: 10.4067/S0716-02082004000100003] - Reguero, M. A., Dozo, M. T. & Cerdeño, E., 2007: A poorly known rodentlike mammal (Pachyrukhinae, Hegetotheriidae, Notoungulata) from the Deseadan (Late Oligocene) of Argentina. Paleoecology, biogeography, and radiation of the rodentlike ungulate in South America.
–Journal of Paleontology: Vol. 81, #6, pp. 1301-1307 [doi: 0022-3360/07/0081-1301] - Reguero, M. A., Ubilla, M. & Perea, D., 2003: A new species of Eopachyrucos (Mammalia; Notoungulata, Interatheriidae) from the Late Oligocene of Uruguay.
–Journal of Vertebrate Paleontology: Vol. 23, #2, pp. 445-457