After Polly, 1996, McKenna & Bell, 1997, Gunnell, 1998, Egi, Tsubamoto & Takai, 2007, and Peigné, Morlo, Chaimanee, Ducrocq, Tun & Jaeger, 2007

<==o †Hyeanodontidae Leidy, 1869 sensu lato [Hyaenodontoidea Trouessart, 1885; Hyaenodontida Haeckel, 1895] [incl. Proviverridae Schlosser, 1886; Limnocyonidae Gazin, 1946]
   |?- †Parvagula palulae Lange-Badré in Godinot, Crochet, Hartenberger, Lange-Badré, Russell & Sigé, 1987; E.Eoc. Eu.
   |?- †Tylodon Gervais, 1848 [Tulodon Zittel, 1893] [nomen oblitum] [non Tylodon Banker 1902, a genus of Fungi]
   |?- †Ischnognathus savagei Stovall, 1948
   `==o †Proviverrinae Schlosser, 1886 [Proviverridae Schlosser, 1886; Proviverrida Haeckel, 1895; Proviverrinae Matthew, 1909; Proviverrini Van Valen, 1966] [incl. Stypolophinae Trouessart, 1885] [paraphyletic]
      |?- †Acarictis ryani Gingerich & Deutsch, 1989
      |?- †Alienetherium bruxwilleri Lange-Badre, 1981
      |?- †Boualitomus marocanensis Gheerbrant, Iarochene, Amaghzaz & Bouya, 2006
      |?- †Gazinocyon vulpeculus (Matthew, 1915) Polly, 1996
      |?- †Galecyon mordax (Matthew, 1915) Gingerich & Deutsch, 1989
      |?-o †Lesmesodon edingeri Morlo & Habersetzer, 1999
      |  |-- †L. edingeri (Springhorn, 1982) [Proviverra edingeri Springhorn, 1982]
      |  `-- †L. behnkeae Morlo & Habersetzer, 1999
      |?- †Matthodon tritens Lange-Badré & Haubold, 1990
      |?- †Praecodens acutus Lange-Badre, 1981
      |?- †Protoproviverra palaeonictides (Lemoine, 1880) Lemoine, 1891 [=? Proviverra palaeonictides Lemoine, 1880] [nonProtoproviverra manzaniana Ameghino, 1891] (a non-creodont (Lemoine, 1891))
      |?-o †Propterodon Martin, 1906
      |  |-- †P. morrisi (Matthew & Granger, 1924)
      |  `-- †P. irdinensis Matthew & Granger, 1925
      |?- †Proviverroides piercei Bown, 1982
      |?-o †Pyrocyon Gingerich & Deutsch, 1989
      |  |-- †P. dioctetus Gingerich & Deutsch, 1989
      |  `-- †P. hians
      |?- †Quercitherium tenebrosum Filhol, 1880 emend. McKenna & Bell, 1997 [pro Quercytherium Filhol, 1880]
      |?- †Tinerhodon disputatum Gheerbrant, 1995
      |?-o †Tritemnodon Matthew, 1906
      |  |?- ?†T. strenuus
      |  `-- †T. agilis (Marsh, 1872)
      |--o †Proviverra Rütimeyer, 1862 [Sinopa Leidy, 1871; Prorhyzaena Rütimeyer, 1891; Leonhardtina Matthes, 1952; Geiselotherium Matthes, 1952]
      |  |-- †P. typica Rütimeyer, 1862
      |  `-- †P. minor (Filhol, 1877)
      `--+--+-- †Leonhardtina gracilis Matthes, 1952
         |  `--o †Allopterodon Ginsburg in Ginsburg et al., 1977 [Hurzelerius Lange-Badré, 1984]
         |     |-- †A. phonax (Van Valen, 1965)
         |     |-- †A. theriodis (Van Valen, 1965
         |     `-- †A. torvidus
         `--+--+-- †Prodissopsalis eocaenicus Matthes, 1952 [Imperatoria Matthes, 1952]
            |  `--o †Eurotherium Polly & Lange-Badré, 1993
            |     |-- †E. theriodis (Van Valen, 1965)
            |     `-- †E. matthesi (Lange-Badré & Haubold, 1990)
            `--+--+-- †Paracynohyaenodon schlosseri Martin, 1906
               |  `--o †Cynohyaenodon Filhol, 1873 [Pseudosinopa Depéret, 1917]
               |     |-- †C. cayluxi Filhol, 1873
               |     |-- †C. trux Van Valen, 1965
               |     `-- †C. lautricensis Lange-Badré, 1978
               `--+--+--o †Arfia Van Valen, 1965
                  |  |  |-- †A. opisthotoma (Matthew, 1901)
                  |  |  |-- †A. shoshoniensis (Matthew, 1915)
                  |  |  |-- †A. zele Gingerich & Deutsch, 1989
                  |  |  |-- †A. junnei Gingerich, 1989
                  |  |  `-- †A. langebadreae
                  |  `--+--o “African †Sinopa”
                  |     |  |-- †“Sinopaatrox 
                  |     |  |-- †“Sinopaaethiopica
                  |     |  `-- †“Sinopaephalmos
                  |     `--+-- †Hyaninailourinae Pilgrim, 1932 sensu Peigné et al., 2007 [Hyaenaelurinae Pilgrim, 1932; Hyaninailourini Ginsburg, 1980; Pterodontinae Polly, 1996; Hyaenaelurinae Egi et al., 2007; Hyaninailourinae Peigné et al., 2007]
                  |        `--+--o †Paratritemnodon Ranga Rao, 1973
                  |           |  |-- †P. jandewalensis Thewissen, Williams & Hussain, 2001
                  |           |  `-- †P. indicus Ranga Rao, 1973
                  |           `--+?- †Yarshea cruenta Egi, Holroyd, Tsubamoto, Shigehara, Takai & Tun, 2004
                  |              `-- †Kyawdawia lupina Egi, Holroyd, Aung Naing Soe, Takai & Ciochon, 2005
                  `--+?-o †Masrasector Simons & Gingerich, 1974
                     |  |-- †M. pithecodacos
                     |  |-- †M. ligabetti
                     |  |-- †M. aegypticum Simons & Gingerich, 1974
                     |  `-- †M. ligabuei Crochet, Thomas, Rogers, Sen & Al-Sulaimani, 1990
                     |--+--o †Prototomus Cope, 1874 [Prolimnocyon] [treated as Limnocyonines by Morlo & Gunnell, 2005]
                     |  |  |-- †P. viverrinus Cope, 1874
                     |  |  |-- †P. secundarius Cope, 1875
                     |  |  |-- †P. multicuspus Cope, 1875
                     |  |  |-- †P. robustus (Matthew, 1915)
                     |  |  |-- †P. deimos Gingerich & Deutsch, 1989
                     |  |  |-- †P. phobos Gingerich & Deutsch, 1989
                     |  |  `-- †P. martis Gingerich & Deutsch, 1989
                     |  |--o †Sinopa Leidy, 1871b [Stypolophus Cope, 1872; ?Triacodon]
                     |  |  |-- †S. rapax Leidy, 1871b [Sinopa minor Wortman, 1902; Stypolophus aculeatus; Triacodon fallax]
                     |  |  |-- †S. pungens (Cope, 1872b) [Stypolophus pungens Cope, 1872]
                     |  |  `-- †S. major Wortman, 1902 [incl.S. grangeri Matthew, 1906]
                     |  `== other “Prototomus-like” Proviverrinae
                     `--+-- †Limnocyoninae Wortman, 1902 sensu McKenna & Bell, 1997 [Limnocyonidae Gazin, 1946; Limnocyonini Van Valen, 1966]
                        `?- †Hyeanodontinae Leidy, 1869 sensu Polly, 1996

After Peigné, Morlo, Chaimanee, Ducrocq, Tun & Jaeger, 2007 (Fig. 4B)

<==o †Hyeanodontidae
   `==o †Proviverrinae
      |== Proviverra-like” Proviverrinae
      `--+--+-- †Arfia
         |  |== Afria-like” South Asian Proviverrinae
         |  `--+== Dissopsalis-like” Proviverrinae
         |     `==Hyaninailourinae
         `--+== Prototomus-like” Proviverrinae
            `--+-- †Prolimnocyon
               `-- †Limnocyoninae

Reference(s):

  • Peigné, S., Morlo, M., Chaimanee, Y., Ducrocq, S., Tun, S. T. & Jaeger, J.-J., 2007: New discoveries of hyaenodontids (Creodonta, Mammalia) from the Pondaung Formation, middle Eocene, Myanmar - paleobiogeographic implications.
    –Geodiversitas: Vol. 29, #3, pp. 441-458

After McKenna & Bell, 1997

<==o †Hyaenodontidae
   |-- †Parvagula
   |?- †Tylodon
   |--o †Limnocyoninae
   |  |--o †Limnocyonini
   |  |  |-- †Prolimnocyon
   |  |  |-- †Thinocyon
   |  |  |-- †Limnocyon
   |  |  |-- †Oxyaenodon
   |  |  |-- †Prolaena
   |  |  `-- †Thereutherium
   |  `--o †Mechaeroidini
   |     |-- †Apataelurus
   |     `-- †Machaeroides
   `--o †Hyeanodontinae
      |--o †Proviverrini
      |  |-- †Prototomus
      |  |-- †Arfia
      |  |-- †Sinopa
      |  |-- †Gazinocyon
      |  |-- †Tritemnodon
      |  |-- †Acarictis
      |  |-- †Protoproviverra
      |  |-- †Galecyon
      |  |-- †Proviverra
      |  |-- †Prodissopsalis
      |  |-- †Paratritemnodon
      |  |-- †Pyrocyon
      |  |-- †Propterodon
      |  |-- †Cynohyaenodon
      |  |-- †Quercitherium
      |  |-- †Praecodens
      |  |-- †Matthodon
      |  |-- †Proviverroides
      |  |-- †Paracynohyaenodon
      |  |-- †Alienetherium
      |  |-- †Allopterodon
      |  |-- †Eurotherium
      |  |-- †Masrasector
      |  |-- †Metasinopa
      |  |-- †Anasinopa
      |  `-- †Dissopsalis
      |--o †Hyaenodontini
      |  |-- †Oxyaenoides
      |  |-- †Francotherium
      |  |-- †Pterodon
      |  |-- †Hyaenodon
      |  |-- †Hemipsalodon
      |  |-- †Consobrinus
      |  |-- †Paenoxyaenoides
      |  |-- †Parapterodon
      |  |-- †Ischnognathus
      |  |-- †Hyainailouros
      |  |-- †Leakitherium
      |  `-- †Megistotherium
      |--o †Teratodontini
      |  `-- †Teratodon
      `--o †Apterodontini
         `-- †Apterodon

After Polly, 1996

<==o †Hyeanodontidae Leidy, 1869 sensu lato
   `==o †Proviverrinae
      |--+-- †Limnocyoninae
      |  `--+-- †Arfia
      |     `--+-- †Dissopsalis
      |        `-- †Pterodontinae
      `--o †Hyeanodontidae sensu stricto?
         |?- †Prototomus
         `--+-- †Sinopa
            `--+-- †Gazinocyon
               `--+-- †Eurotherium
                  `--+-- †Propterodon
                     `-- †Hyeanodontinae

Reference(s):

  • Polly, P. D., 1996: The skeleton of Gazinocyon vulpeculus gen. et comb. nov. and the cladistic relationships of Hyaenodontidae (Eutheria, Mammalia).
    –Journal of Vertebrate Paleontology: Vol. 16, #2, pp. 303-319

Reference(s):