After McKenna & Bell, 1997, Gunnell & Bartels, 1999, Zonneveld, Gunnell & Bartels, 2000, and Scott, Fox & Youzwyshyn, 2002, with species from Alroy, 2002
<==o †Pantolestidae Cope, 1884 [taxonomic history] |?- †Apheliscus [is an apheliscid ungulate] |?-o †Paleotomus Van Valen, 1967 [taxonomic history] [treated as cimolestid by McKenna & Bell, 1997] | |-- †P. junior Scott, Fox & Youzwyshyn, 2002 | |-- †P. senior (Simpson, 1937) [taxonomic history] | |-- †P. carbonensis Secord, 1998 | |-- †P. radagasti (Van Valen, 1978) [taxonomic history] | `-- †P. milleri Rigby, 1980 |--o †Pentacodontinae Simpson, 1937 [taxonomic history] | |?- †Protentomodon ursirivalis Simpson, 1928a; U.Paleoc. NA. [treated as cimolestid by McKenna & Bell, 1997] | |--o †Aphronorus Simpson, 1935 [Aphronomus (lapsus calami)]; L.-U.Paleoc. NA. | | |-- †A. fraudator Simpson, 1935d | | |-- †A. orieli Gazin, 1969 | | |-- †A. ratatoski Winterfeld, 1982 | | `-- †A. simpsoni Gazin, 1938 | |--o †Bisonalveus Gazin, 1956; U.Paleoc. NA. | | |-- †B. browni Gazin, 1956 | | `-- †B. holtzmani Gingerich, 1983c | |--o †Coriphagus Douglass, 1908 [†Pantomimus Van Valen, 1967] [incl. †Mixoclaenus Matthew & Granger, 1921] L.Paleoc. NA. | | |-- †C. montanus Douglass, 1908 | | `-- †C. encinensis (Matthew & Granger, 1921) [†Mixoclaenus encinensis Matthew & Granger, 1921); †Coriphagus encinensis Simpson, 1937a, Williamson & Lucas, 1993, Williamson, 1996, and Alroy, 2002] | |--o †Pentacodon Scott, 1892; L.Paleoc. NA. | | |-- †P. inversus (Cope, 1888) [†Chriacus inversus Cope, 1888; †Pentacodon inversus Scott, 1892, Matthew, 1937, Williamson & Lucas, 1993, Williamson, 1996, and Alroy, 2002] | | `-- †P. occultus Matthew, 1937 | `-- †Amaramnis gregoryi Gazin, 1962; L.Eoc. NA. |--o Pantolestinae Cope, 1884 [taxonomic history] | |?- †Propalaeosinopa albertensis Simpson, 1927; M.Paleoc. NA. (nomen dubium, invalid sr. synon. of †Bessoecetor) | |-- †Bessoecetor septentrionalis (Russell, 1929) Simpson, 1936 sensu Scott, Fox & Youzwyshyn, 2002; L.-U.Paleoc. NA. ?U.Paleoc. Eu. [Synonyms] | |--o †Palaeosinopa Matthew, 1901 [taxonomic history] | | |-- †P. dorri Gingerich, 1980c | | |-- †P. lutreola Matthew, 1918b [non 1899] | | |-- †P. incerta Bown & Schankler, 1982 | | |-- †P. simpsoni Van Valen, 1967 [taxonomic history] | | `--+-- †P. didelphoides (Cope, 1881) [taxonomic history] | | `-- †P. veterrima Matthew, 1901 | |-- †Thelysia artemia Gingerich 1982; U.Paleoc. NA. [a pantolestid?] | |-- †Pagonomus dionysi D.E. Russell, 1964; U.Paleoc. NA. | |-- †Todralestes variabilis Gheerbrant, 1991; U.Paleoc.-L.Eoc. Af. | |--o †Pantolestes Cope, 1872b [incl.? Passalacodon Marsh, 1872; †Anisacodon Marsh, 1872] M.Eoc. EAs. NA. | | |-- †P. longicaudus Cope, 1872 [†Pantolestes longicaudus Cope, 1872; †Tomitherium longicaudum (Cope); Cope, 1873; †Passalacodon littoralis Marsh, 1872; †Pantolestes longicaudus Cope; Matthew, 1909, Gazin, 1958a, Gazin, 1976, and Alroy, 2002) | | |-- †P. elegans (Marsh, 1872) | | |-- †P. natans Matthew, 1909b | | |-- †P. phocipes Matthew, 1909b | | `-- †P. intermedius Matthew, 1909b | |-- †Bogdia Dashzeveg & D.E. Russell, 1985; M.Eoc. EAs. | |-- †Chadronia margaretae Cook, 1954 [incl. †Cymaprimadon J. Clark, 1968] U.Eoc. ?As. NA. | |--o †Buxolestes Jaeger, 1970; M.Eoc. Eu. | | |-- †P. piscator | | `-- †P. minor | |-- †Bouffinomus Mathis, 1989; U.Eoc. Eu. | |-- †Galethylax blainvillei Gervais, 1850; U.Eoc. Eu. | `-- †Oboia argillaceus Gabunia, 1989; L.Olig. As. [also treated as merialine paroxyclaenid] `--o †Dyspterninae Kretzoi, 1943 [taxonomic history] |-- †Fordonia Marandat, 1989; L.Eoc. Eu. [non Fordonia Gray, 1842(?) (Serpentes: Colubridae: Homalopsinae)] |--o †Cryptopithecus Schlosser, 1890 [incl. †Opsiclaenodon Butler, 1946; †Androconus Quinet, 1965] U.Eoc.-L.Olig. Eu. | |-- †C. siderolithicus | `-- †C. macrognathus |--o †Dyspterna Hopwood, 1927; U.Eoc.-L.Olig. Eu. | |-- †D. woodi | `-- †D. helbingi Dal Piaz, 1930 |--o †Kochictis Kretzoi, 1943; U.Olig. Eu. | `-- †K. centenii `-- †Gobipithecus Dashzeveg & D.E. Russell, 1992; L.Olig. As.
Reference(s):
- Alroy, J., 2002: North American fossil mammal systematics database.
–iNet: North American fossil mammal systematics database: http://www.nceas.ucsb.edu/~alroy/nafmsd.html - Burkitt, J. H., 1995: Mammals: A world listing of living and extinct species.
–iNet: [no longer available, looking for new link] - Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698. - Carroll, R. L., 1988: Appendix. 594-648.
in Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698. - Fox, R. C. & Scott, C. S., 2005: First evidence of a venom delivery apparatus in extinct mammals.
–Nature: Vol. 435, #7045, pp. 1091-1093 [doi: 10.1038/nature03646] - Gazin, C. L., 1976: Mammalian faunal zones of the Bridger Middle Eocene.
–Smithsonian Contributions to Paleobiology: No. 26, pp. iv-25 - Gunnell, G. F. & Bartels, W. S., 1999: Middle Eocene vertebrates from the Uinta Basin, Utah, and their relationship with faunas from the southern Green River Basin, Wyoming. 429-442
in Gillette, D. D., (eds.) 1999: Vertebrate Paleontology in Utah.
–Utah Geological Survey, Salt Lake City, 1999 - McKenna, M. C. & Bell, S. K., (eds.) 1997: Classification of mammals – above the species level.
–Columbia University Press, New York, 1997, xii-631 - Scott, C. S., Fox, R. C. & Youzwyshyn, G. P., 2002: New earliest Tiffanian (Late Paleocene) mammals from Cochrane 2, southwestern Alberta, Canada.
–Acta Palaeontologica Polonica: Vol. 47, #4, pp. 691-704 - Zonneveld, J.-P., Gunnell, G. F. & Bartels, W. S., 2000: Early Eocene fossil vertebrates from the southwestern Green River basin, Lincoln and Uinta Counties, Wyoming.
–Journal of Vertebrate Paleontology: Vol. 20, #2, pp. 369-386