After McKenna & Bell, 1997, and Eberle, 1999, with species from Alroy, 2002, and other unspecified sources
Alternative phylogeny after Alroy, 2002
<==o †Didelphodonta McKenna, 1995 `--o †Cimolestidae Marsh, 1889 [taxonomic history] |?-o †Aboletylestes Russell, 1964; L.Pal. Af. Eu. [treated as cimolestid by McKenna & Bell, 1997] | |-- †A. hypselus Russell 1964 | `-- †A. robustus Gheerbrant, 1992 |?- †Betonnia tsosia Williamson, Weil & Standhardt, 2011 [“Cimolestes sp.” Standhardt, 1980; “Prodiacodon estevani” Standhardt, 1980]; E.Pal. SWNAm. |?- †Chacopterygus minutus Williamson, Weil & Standhardt, 2011 [“Procerberus sp.” Standhardt, 1980; “Procerberus? sp.” Standhardt, 1980]; E.Pal. SWNAm. |--+-- †Batodon tenuis Marsh, 1892 [is a soricomorph??] | `-- †Maelestes gobiensis Wible, Rougier, Novacek & Asher, 2007, L.Cret. EAs. |?-+--o †Didelphodus Cope, 1882b [Didelphyodus, Phenacops] [treated as paleoryctid] | | |--+-- †D. absarokae | | | `-- †D. altidens (Marsh, 1872) | | `--+-- †D. rheos | | `-- †D. serus | `--+-- †Avunculus didephodonti Van Valen, 1966; L.Pal. NA. [treated as paleoryctid] | `--+-- †Acmeodon secans Matthew & Granger, 1921; E-L.Pal. NA. [treated as paleoryctid] | `--o †Gelastops Simpson, 1935d [†Emperodon Simpson, 1935d]; E.-L.Pal. NA. [treated as paleoryctid] | |-- †G. joni Rigby, Jr., 1980 | `-- †G. parcus Simpson, 1935d [†Emperodon acmeodontoides Simpson, 1935d; Gelastops parcus Simpson; Simpson,1937a, Van Valen,1966] |?-o †Paleotomus Van Valen, 1967; L.Pal. NA. [recently treated as a pantolestid] | |-- †P. junior Scott, Fox & Youzwyshyn, 2002 | |-- †P. senior (Simpson, 1937) [taxonomic history] | |-- †P. carbonensis Secord, 1998 | |-- †P. radagasti (Van Valen, 1978) [taxonomic history] | `-- †P. milleri Rigby, 1980 |?- †Tinerhodon disputatum Gheerbrant, 1995; L.Pal. ?Eu. NWAf. |?- †Protentomodon ursirivalis Simpson, 1928a; L.Pal. NA. [treated as pantolestid by Alroy (2002)] |?- †Ilerdoryctes sigei Marandat, 1989; L.Eoc. Eu. |?- †Tsaganius ambiguus D.E. Russell & Dashzeveg, 1986; E.Eoc. As. [treated as paleoryctid] `--+?-o †Gallolestes Lillegraven, 1976 [is a zhelestid?? Also listed as a Leptictida incertae sedis by McKenna & Bell, 1997] | |-- †G. pachymandibularis Lillegraven, 1976 | `-- †G. agujaensis Cifelli, 1994 `--+==o †Procerberus Sloan & Van Valen, 1965; E.Pal. NA. | |?- †P. andetiticus Middleton & Dewar, 2004 | |?- †P. plutonis Van Valen, 1978 | |-- †P. formicarum Sloan & Van Valen, 1965 | `--+-- †P. grandis Middleton & Dewar, 2004 | `--+-- †Alveugena carbonensis Eberle, 1999; E.Pal. NA. | `-- †Taeniodonta Cope, 1876 `--+?- †Telacodon laevis Marsh, 1892 |?- †Puercolestes simpsoni Reynolds, 1936 [“Genus B, Puerco” Van Valen, 1966; Cimolestes simpsoni (Reynolds); Clemens, 1973; “Cimolestes sp.” Standhardt, 1980; “Procerberus sp.” Standhardt, 1980; Puercolestes simpsoni Reynolds; Williamson et al., 2011] |?- †Altacreodus magnus (Clemens & Russell, 1965) Fox, 2015 [Cimolestes magnus Clemens & Russell, 1965] |?- †Ampilestes cerberoides (Lillagraven, 1969) Fox, 2015 [Cimolestes cerberoides Lillagraven, 1969] |?- †Scollardius propaleoryctes (Lillagraven, 1969) Fox, 2015 [Cimolestes propaleoryctes Lillagraven, 1969] `--o †Cimolestes Marsh, 1889 sensu Fox, 2015 [incl. †Nyssodon Simpson, 1927; Puercolestes Reynolds, 1936] L.Cret.-E.Pal. NA., ?E.Pal. SA., L.Pal. Af. |-- †C. incisus Marsh, 1889 [Nyssodon incisus] `-- †C. stirtoni Clemens, 1973
After Alroy, 2002<==o Didelphodonta |?-o ?†Geolabididae | |-- †Batodon | `--+-- †Centetodon | `--+-- †Marsholestes | `-- †Batodontoides `--+--o ?†Didelphodontidae | |-- †Cimolestes incisus | |-- †Cimolestes simpsoni | `--+--o †Didelphodus | | |--+-- †D. absarokae | | | `-- †D. altidens | | `--+-- †D. rheos | | `-- †D. serus | `--+-- †Avunculus | `--+-- †Acmeodon | `-- †Gelastops `--o ?†Cimolestidae |?- †Telacodon |?- †Gallolestes |?-o †Cimolestes | |-- †C. magnus | |-- †C. cerberoides | |-- †C. propaleoryctes | |-- †C. stirtoni | `-- †C. sp. `--+-- †Procerberus `--+-- †Alveugena `-- †Stylinodontidae Reference(s):
|
Reference(s):
- Alroy, J., 2002: North American fossil mammal systematics database.
–iNet: North American fossil mammal systematics database: http://www.nceas.ucsb.edu/~alroy/nafmsd.html - Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698. - Carroll, R. L., 1988: Appendix. 594-648.
in Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698. - Eberle, J. J., 1999: Bridging the transition between didelphodonts and taeniodonts.
–Journal of Paleontology: Vol. 73, No. 5, pp. 936-944 - Fox, R. C., 2015: A revision of the Late Cretaceous–Paleocene eutherian mammal Cimolestes Marsh, 1889.
–Canadian Journal of Earth Sciences: in press [doi: 10.1139/cjes-2015-0113] - McKenna, M. C. & Bell, S. K., (eds.) 1997: Classification of mammals – above the species level.
–Columbia University Press, New York, 1997, xii-631 - Scott, C. S., Fox, R. C. & Youzwyshyn, G. P., 2002: New earliest Tiffanian (Late Paleocene) mammals from Cochrane 2, southwestern Alberta, Canada.
–Acta Palaeontologica Polonica: Vol. 47, #4, pp. 691-704 - Wible, J. R., Rougier, G. W., Novacek, M. J. & Asher, R. J., 2007: Cretaceous eutherians and Laurasian origin for placental mammals near the K/T boundary.
–Nature: Vol. 447, #7147, pp. 1003-1006 [doi: 10.1038/nature05854] - Williamson, T. E., Weil, A. & Standhardt, B., 2011: Cimolestids (Mammalia) from the Early Paleocene (Puercan) of New Mexico.
–Journal of Vertebrate Paleontology: Vol. 31, #1, pp. 162-180 [doi: 10.1080/02724634.2011.539649]