After Bergmans, 1997 [from Giannini & Simmons, 2003]
<==o Cynopterinae Andersen, 1912
|--o Cynopterus ()?
| |-- C. brachyotis ()?
| |-- C. minor ()?
| |-- C. archipelagus ()?
| |-- C. horsfieldi ()?
| |-- C. sphinx (lyhytkuonohedelmälepakko)
| |-- C. titthaecheilus ()?
| `-- ?C. nusatenggara ()?
|--o Megaerops ()?
| |-- M. niphanae ()?
| |-- M. ecaudatus ()?
| |-- M. kusnotoi ()?
| `-- M. wetmorei ()?
|--o Ptenochirus ()?
| |-- P. minor ()?
| `-- P. jagori ()?
|-- Dyacopterus spadiceus ()?
|-- Chironax melanocephalus (hattuhedelmälepakko)
|--o Thoopterus nigrescens ()?
| |-- T. n. nigrescens ()?
| `-- T. n. latidens ()?
|-- Sphaerias blanfordi ()?
|-- Balionycteris maculata (täpläpoimuhedelmälepakko)
|-- Aethalops alecto (kääpiöhedelmälepakko)
|-- Penthetor lucasi ()?
|-- Latidens salimalii ()?
|-- Alionycteris paucidentata ()?
|-- Otopteropus cartilagonodus ()?
`-- Haplonycteris fischeri ()?
Reference(s):
- Butler, P. C., 1978: Insectivora and Chiroptera. 56-68
in Maglio, V. J. & Cooke, H. B. S., (eds.) 1978: Evolution of African Mammals, 1.
–Harvard University Press, Cambridge, Massachusetts, London, England, 1978, xiv-641
- Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698.
- Carroll, R. L., 1988: Appendix. 594-648.
in Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698.
- Giannini, N. P. & Simmons, N. B., 2003: A phylogeny of megachiropteran bats (Mammalia: Chiroptera: Pteropodidae) based on direct optimization analysis of one nuclear and four mitochondrial genes.
–Cladistics: Vol. 19, pp. 496-511
- McKenna, M. C. & Bell, S. K., (eds.) 1997: Classification of mammals – above the species level.
–Columbia University Press, New York, 1997, xii-631
- Nowak, R. M., 1991: Walker's Mammals of the World; Part 1.
–The Johns Hopkins University Press, Baltimore and London, 1991, xlviii-642-lxiii
- Nowak, R. M., 1991: Walker's mammals of the World; Part 2.
–The Johns Hopkins University Press, Baltimore and London, 1991, xii-643-1629