After Lambert, 2005, Bianucci, Lambert & Post, 2007, and Lambert, de Muizon & Bianucci, 2013, with fossil genera from McKenna & Bell, 1997 and Recent species from Nowak, 1990
<==o Hyperoodontoidea Gray, 1846 sensu McKenna & Bell, 1997 [Ziphioidea Gray, 1850] [taxonomic history] (nokkavalasmaiset hammasvalaat) |?- †Palaeoziphius scaldensis Abel, 1905 [Odontoceti incertae sedis sensu Lambert, 2005] |?- †Squaloziphius emlongi de Muizon, 1991; L.Mioc. ENAm. |--+--o †Messapicetus Bianucci, Landini & Varola, 1992; U.Mioc. SEu. | | |-- †M. longirostris Bianucci, Landini & Varola, 1992 | | `-- †M. gregarius Bianucci, Lambert & Post, 2010 | `--+-- †Beneziphus brevirostris Lambert, 2005; ?U.Mioc.-?L.Plioc. WEu. | `--o †Ziphirostrum du Bus, 1868 [taxonomic history] | |-- †Z. marginatum du Bus, 1868 sensu Lambert, 2005 [taxonomic history] | |-- †Z. turniense du Bus, 1868 [taxonomic history] | `-- †Z. recurvus (du Bus, 1868) sensu Lambert, 2005 [taxonomic history] `--o Hyperoodontidae Gray, 1846 sensu McKenna & Bell, 1997 [Ziphiidae Gray, 1850; †Squalodelphidae Dal Piaz, 1916] [taxonomic history] (beaked whales; nokkavalaat) |--o Berardiinae Moore, 1968 sensu Lambert, 2005 (giant beaked whales; berardinvalaat) | |?- †Archaeoziphius microglenoideus Lambert & Louwye, 2006 [Mioziphius belgicus Abel, 1905 (in partim)] M.Mioc. WEu. | |?- †Microberardius africanus Bianucci, Lambert & Post, 2007 | `--o Berardius Duvernoy, 1851 (giant beaked whales; berardinvalaat) | |-- B. arnuxii Duvernoy, 1851 (Arnoux's beaked whale; etelänberardinvalas) | `-- B. bairdii Stejneger, 1883 (Baird's beaked whale; berardinvalas) `--+-- †Nenga meganasalis Bianucci, Lambert & Post, 2007 |-- Tasmacetus shepherdi Oliver, 1937 [Tasmacetina Moore, 1968] (tasmanianvalas) [has been treated as Berardiine] |--+-- †Xhosacetus hendeysi Bianucci, Lambert & Post, 2007 | `--+-- †Pterocetus benguelae Bianucci, Lambert & Post, 2007 | `--o Hyperoodontinae Gray, 1846 sensu Lambert, 2005; Bianucci, Lambert & Post, 2007 | |?- †Khoikhoicetus agulhasis Bianucci, Lambert & Post, 2007 | |-- Indopacetus pacificus Moore, 1968 [Indocetina Moore, 1968] (Tropical bottlenose whale, Longman's beaked whale; longmaninvalas) | `--+-- †Africanacetus ceratopsis Bianucci, Lambert & Post, 2007 | |-- †Ihlengesi saldanhae Bianucci, Lambert & Post, 2007 | `==o Mesoplodon Gervais, 1850 | |?- †M. longirostris | |?- M. perrini Dalebout, Mead, Baker, Baker & van Helden, 2002 (Perrin's beaked whale; kaliforniannokkavalas) | |?- M. peruvianus Reyes, Mead & Van Waerebeek, 1991 (Pygmy beaked whale) | |==o (Mesoplodon) | | |?- M. (M.) hectori Gray, 1871 (Hector's beaked whale; hektorinvalas) | | |?- M. (M.) europaeus (Gervais, 1855) (Gervais' beaked whale; gervaisinvalas) | | |?- M. (M.) ginkgodens Nishiwaki & Kamiya, 1958 (Ginkgo-toothed beaked whale; ginkgohammasvalas) | | |?- M. (M.) carlhubbsi Moore, 1963 (Hubbs' beaked whale; hubbinvalas) | | |?- M. (M.) bowdoini Andrews, 1908 (Andrews' beaked whale; andrewinvalas) | | |?- M. (M.) stejnegeri True, 1885 (Stejneger's beaked whale; stejnegerinvalas) | | |-- M. (M.) bidens Sowerby, 1804 (Sowerby's beaked whale; kaksihammasvalas, sowerbynvalas) | | `--+-- †M. slangkopi Bianucci, Lambert & Post, 2007 | | |-- M. (M.) grayi von Haast, 1876 (Gray's beaked whale; graynvalas) | | `--+-- M. (Dioplodon) densirostris Blainville, 1817 (Blainville's beaked whale; blainvillenvalas) | | `--o (Dolichodon) | | |?- M. (D.) traversii (Gray, 1874) (Spade-toothed whale) | | `-- M. (D.) layardii Gray, 1865 (Strap-toothed whale; kuonokoppavalas) | `--+?- Mesoplodon (Mesoplodon) mirus True, 1913 (True's beaked whale; truenvalas) | `--o Hyperoodon Lacépède, 1804 (bottlenose whales) | |-- H. ampullatus (Forster, 1770) (Northern bottlenose whale; nokkavalas, pohjannokkavalas) | `-- H. planifrons (Flower, 1882) (Southern bottlenose whale; etelännokkavalas) `--o Ziphiinae Gray, 1850 sensu Lambert, 2005 |?-o †Aporotus du Bus, 1868 | |-- †A. recurvirostris du Bus, 1868 | `-- †A. dicyrtus du Bus, 1868 |--+--o †Izikoziphius Bianucci, Lambert & Post, 2007 | | |-- †I. rossi Bianucci, Lambert & Post, 2007 | | `-- †I. angustus Bianucci, Lambert & Post, 2007 | `--o Ziphius Cuvier, 1823 | |-- †Z. sp. [Bianucci, Lambert & Post, 2007] | |-- †Z. compressus Owen 1870 | `-- Z. cavirostris Cuvier, 1823 (cuvierinvalas) `--+--o †Choneziphius Duvernoy, 1851 [Proroziphius Leidy, 1876] | |?- †C. chonops (Leidy, 1876) Trouessart, 1898 [Proroziphius chonops Leidy, 1876] | |-- †C. leidyi Bianucci, Miján, Lambert, Post & Mateus, 2013 | |?- †C. liops Leidy, 1876 | |-- †C. planirostris (Cuvier, 1823) [Ziphius planirostris Cuvier, 1823; Ziphius cuvieri Owen, 1870; Ziphius planus; Choneziphius packardi Lankester, 1870; Choneziphius trachops Leidy 1876 (incertae sedis)] | `?- †C. macrops (Leidy, 1876) [Proroziphius macrops Leidy, 1876] `--+-- †Imocetus piscatus Bianucci, Miján, Lambert, Post & Mateus, 2013 `--+-- †Globicetus hiberus Bianucci, Miján, Lambert, Post & Mateus, 2013 `--o †Tusciziphius Bianucci, 1997; ?M.Mioc.-Plioc. SEu. |-- †T. crispus Bianucci, 1997 `-- †T. atlanticus Bianucci, Miján, Lambert, Post & Mateus, 2013
Hyperoodontidae not treated by Lambert, 2005
†Belemnoziphius Huxley, 1864
†B. compressus Huxley, 1864
†B. prorops Leidy, 1876
?†Caviziphius altirostris Bianucci & Post, 2005; ?L.Mioc. WEu.
†Pelycorhamphus pertortus Cope, 1895 [nomen dubium]
†Cetorhynchus christoli (Gervais, 1861) [Mesoplodon (Cetorhynchus) christoli Gervais, 1861]
†Nazcacetus urbinai Lambert, Bianucci & Post, 2009
?†Ziphioides Probst, 1886 (nomen dubium)
†Anoplonassa forcipata Cope, 1869
†Eboroziphius coelops Leidy, 1877
†Ninoziphius platyrostris de Muizon, 1983; L.Plioc. WSAm.
†Berardiopsis pliocaenus Portis, 1886
After Lambert, de Muizon & Bianucci, 2013<==o Ziphiidae sensu lato |-- †Ninoziphius `--+--+-- †Messapicetus | `--+-- †Beneziphus | `-- †Ziphirostrum `--o Ziphiidae sensu stricto |-- Tasmacetus `--+-- †Nazcacetus |--o Berardiinae | |-- †Archaeoziphius | |-- †Microberardius | `-- Berardius `--+--+-- †Xhosacetus | `--+-- †Pterocetus | `--o Hyperoodontinae | |-- Indopacetus | `--+-- Mesoplodon | `--+-- †Africanacetus | `--+-- †Ihlengesi | `-- Hyperoodon `--o Ziphiinae |--+-- †Izikoziphius | `-- Ziphius `--+-- †Choneziphius `--+-- †Imocetus `--+-- †Tusciziphius `-- †Globicetus Reference(s):
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After Bianucci, Miján, Lambert, Post & Mateus, 2013<==o Ziphiidae sensu lato |--+-- †Messapicetus | `--+-- †Beneziphus | `-- †Ziphirostrum `--o Ziphiidae sensu stricto |-- Tasmacetus |-- †Nazcacetus |--o Berardiinae | |-- †Archaeoziphius | |-- †Microberardius | `-- Berardius |--+-- †Xhosacetus | |-- †Pterocetus | `--o Hyperoodontinae | |-- Indopacetus | `--+-- Mesoplodon | `--+-- †Africanacetus | |-- †Ihlengesi | `-- Hyperoodon `--o Ziphiinae |-- †Imocetus `--+-- †Globicetus `--+== †Tusciziphius `--+-- †Choneziphius `--+-- †Izikoziphius `-- Ziphius Reference(s):
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After Bianucci, Lambert & Post, 2010<==o Ziphiidae sensu lato |--+-- †Messapicetus | |-- †Beneziphus | `-- †Ziphirostrum `--o Ziphiidae sensu stricto |-- Tasmacetus `--+-- †Nazcacetus |--o Berardiinae | |-- †Archaeoziphius | |-- †Microberardius | `-- Berardius |--+-- †Xhosacetus | `--+-- †Pterocetus | `--o Hyperoodontinae | |-- Indopacetus | `--+-- Mesoplodon | |-- †Africanacetus | |-- †Ihlengesi | `-- Hyperoodon `--o Ziphiinae |-- †Tusciziphius `--+-- †Choneziphius `--+-- †Izikoziphius `-- Ziphius Reference(s):
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After Lambert, 2005<==o Ziphiidae sensu lato |-- †Squaloziphius `--o Ziphiidae sensu stricto |-- Tasmacetus |-- Berardius `--+-- Indopacetus |--+-- Mesoplodon | `-- Hyperoodon `--+--+-- †Beneziphus | `-- †Ziphirostrum `--+-- †Choneziphius `--+-- †Tusciziphius `-- Ziphius Reference(s):
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After McKenna & Bell, 1997
Hyperoodontoidea
Hyperoodontidae
†Notocetus
†Medocinia
†Belemnoziphius
†Pelycorhamphus
†Squalodelphis
†Choneziphius
Mesoplodon
†Squaloziphius
†Cetorhynchus
†Ziphirostrum
†Messapicetus
†Palaeoziphius
†Ziphioides
†Anoplonassa
†Proroziphius
†Ninoziphius
Hyperoodon
Ziphius
†Berardiopsis
Berardius
Tasmacetus
Indopacetus
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Reference(s):
- Bianucci, G., Lambert, O. & Post, K., 2010: High concentration of long-snouted beaked whales (genus Messapicetus) from the Miocene of Peru.
–Palaeontology: Vol. 53, #5, pp. 1077-1098 [doi: 10.1111/j.1475-4983.2010.00995.x] - Bianucci, G., Lambert, O. & Post, K., 2007: A high diversity in fossil beaked whales (Mammalia, Odontoceti, Ziphiidae) recovered by trawling from the sea floor off South Africa.
–Geodiversitas: Vol. 29, #4, pp. 561-618 - Bianucci, G., Miján, I., Lambert, O., Post, K. & Mateus, O., 2013: Bizarre fossil beaked whales (Odontoceti, Ziphiidae) fished from the Atlantic Ocean floor of the Iberian Peninsula.
–Geodiversitas: Vol. 35, #1, pp. 105-153 [doi: 10.5252/g2013n1a6] - Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698. - Carroll, R. L., 1988: Appendix. 594-648.
in Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698. - Lambert, O., 2005: Systematics and phylogeny of the fossil beaked whales Ziphirostrum du Bus, 1868 and Choneziphius Duvernoy, 1851 (Mammalia, Cetacea, Odontoceti), from the Neogene of Antwerp (North of Belgium).
–Geodiversitas: Vol. 27, #3, pp. 443-497 - Lambert, O. & Louwye, S., 2006: Archaeoziphius microglenoideus, a new primitive beaked whale (Mammalia, Cetacea, Odontoceti) from the Middle Miocene of Belgium.
–Journal of Vertebrate Paleontology: Vol. 26, #1, pp. 182-191 - Lambert, O., de Muizon, C. & Bianucci, G., 2013: The most basal beaked whale Ninoziphius platyrostris Muizon, 1983: clues on the evolutionary history of the family Ziphiidae (Cetacea: Odontoceti).
–Zoological Journal of the Linnean Society: Vol. 167, #4, pp. 569-598 [doi: 10.1111/zoj.12018] - McKenna, M. C. & Bell, S. K., (eds.) 1997: Classification of mammals – above the species level.
–Columbia University Press, New York, 1997, xii-631 - Nowak, R. M., 1991: Walker's Mammals of the World; Part 1.
–The Johns Hopkins University Press, Baltimore and London, 1991, xlviii-642-lxiii - Nowak, R. M., 1991: Walker's mammals of the World; Part 2.
–The Johns Hopkins University Press, Baltimore and London, 1991, xii-643-1629