After Uhen & Gingerich, 2001
<==o Cetacea Brisson, 1762 [paraphyletic Archaeocete Flower, 1883; Archaeoceti Flower, 1883] (whales; valaat) |--o Pakicetidae Gingerich & Russell, 1990 | |-- †Ichthyolestes pinfoldi Dehm & Oettingen-Spielberg, 1958 | |-- †Nalacetus ratimitus Thewissen & Hussain, 1998 | `--o †Pakicetus Gingerich & Russell, 1981 | |-- †P. attocki (West, 1980) | `-- †P. inachus Gingerich & Russell, 1981 `--+?- †Himalayacetus subathuensis Bajpai & Gingerich, 1988 |?- †Gandakasia potens Dehm & Oettingen-Spielberg, 1958 |--+-- †Ambulocetus natans Thewissen, Madar & Hussain, 1996 [Ambulocetidae Thewissen, Madar & Hussain, 1996] | `--o †Remingtonocetidae Kumar & Sahni, 1986 | |-- †Attockicetus praecursor Thewissen & Hussain, 2000 | |?- †Rayanistes afer Bebej, Zalmout, Abed El-Aziz, Antar & Gingerich,, 2016 | `--+-- †Dalanistes ahmedi Gingerich, Arif & Clyde, 1995 | `--+--o †Remingtonocetus Kumar & Sahni, 1986 | | |-- †R. harudiensis (Sahni & Mishra, 1975) [Protocetus harudiensis Sahni & Mishra, 1975] | | `-- †R. domandaensis Gingerich, ul Haq, Khan & Zalmout, 2001 | `--o †Andrewsiphius Sahni & Mishra, 1975 [taxonomic history] | |-- †A. minor (Sahni & Mishra, 1975) [taxonomic history] | `-- †A. sloani (Sahni & Mishra, 1972) Gingerich, ul Haq, Khan & Zalmout, 2001 [taxonomic history] `==o Protocetidae Stromer, 1908 sensu lato [a paraphyletic assemblage] |?- †Artiocetus clavis Gingerich, ul Haq, Zalmout, Khan & Malkani, 2001 |?- †Eocetus schweinfurthi (Fraas, 1904a) Fraas, 1904b [Mesocetus schweinfurthi Fraas, 1904a] |?- †Maiacetus inuus Gingerich, ul Haq, von Koenigswald, Sanders, Smith & Zalmout, 2009; M.Eoc. Ind. |?- †Natchitochia jonesi Uhen, 1998 |?- †Pappocetus lugardi Andrews, 1920 |?- †Quisracetus arifi Gingerich, ul Haq, Khan & Zalmout, 2001 |?- †Takracetus simus Gingerich, Arif & Clyde, 1995 |--o †Gaviacetus Gingerich, Arif & Clyde, 1995 | |-- †G. razai Gingerich, Arif & Clyde, 1995 | `-- †G. sahnii Bajpai & Thewissen, 1998 `--+--o †Rodhocetus Gingerich et al., 1995a | |-- †R. balochistanensis Gingerich, ul Haq, Zalmout, Khan & Malkani, 2001 | `-- †R. kasrani Gingerich et al., 1995a `--+--o †Babiacetus Trivedy & Satsangi, 1984 | |-- †B. mishrai Bajpai & Thewissen, 1998 | `-- †B. indicus Trivedy & Satsangi, 1984 `--+--o †Protocetidae Stromer, 1908 sensu O'Leary, 1998 [sensu stricto?] | |-- †Indocetus ramani Sahni & Mishra, 1975 [Indocetinae Gingerich et al., 1993] | `-- †Protocetus atavus Fraas, 1904a `--+-- †Georgiacetus vogtlensis Hulbert, Petkewich, Bishop, Bukry & Aleshire, 1998 |?- †Carolinacetus gingerichi Geisler, Sanders & Luo, 2005 `--o Pelagiceti Uhen 2008 `==o †Basilosauridae Cope, 1868 emend. Barnes & Mitchell, 1978 [paraphyletic] |?- †Supayacetus muizoni Uhen, Pyenson, Devries, Urbina & Renne, 2011 |?- †Stromerius nidensis Gingerich, 2007 [Zeuglodon osiris Stromer, 1903 (in partim); Dorudon zitteli Kellogg, 1936 (in partim)] [Stromeriinae Gingerich, 2007] |?- †Masracetus markgrafi Gingerich, 2007 [Zeuglodon isis, Stromer, 1908 (in partim); Zeuglodon cf. brachyspondylus Stromer, 1908 (in partim); Prozeuglodon isis Kellogg, 1936 (in partim); Zeuglodon cfr. brachyspondylus, Kellogg, 1936; Zeuglodon brachyspondylus Slijper, 1936 (in partim)] `--+?-o †Basilotritus Gol'din & Zvonok, 2013 | |-- †B. uheni Gol'din & Zvonok, 2013 | `-- †B. wardii (Uhen, 1999) [“Eocetus” wardii Uhen, 1999] |--o Basilosaurinae Cope, 1868 emend. Barnes & Mitchell, 1978 | |?- †Basiloterus hussaini Gingerich et al., 1997 | `--o †Basilosaurus Harlan, 1834 [Alabamornis Abel, 1906; Hydrarchos Koch, 1845; Hydrarchus Muller, 1849; “Hydrargos” Koch, 1845; “Zeuglodon” Owen, 1839; Zygodon (Owen, 1839)] | |-- †B. cetoides (Owen, 1839) [Alabamornis gigantea Abel, 1906; Hydrarchos sillimani Wyman, 1845; Hydrargos sillimanii Koch, 1845; Zeuglodon ceti Wyman, 1845; Zeuglodon harlani De Kay, 1842; Zeuglodon macrospondylus Müller, 1849] | |-- †B. isis (Andrews, 1904) Gingerich et al. 1990 [Zeuglodon, Prozeuglodon] | `-- †B. drazindai Gingerich et al., 1997 |?-o †Cynthiacetus Uhen, 2005 | |-- †C. maxwelli Uhen, 2005 | `-- †C. peruvianus Martínez-Cáceres & Muizon, 2011 `--o †Dorudontinae Miller, 1923 emend. Barnes & Mitchell, 1978 [paraphyletic] |--o †Dorudon Gibbes, 1845 [Zeuglodon] | |-- †D. atrox (Andrews, 1906) [Prozeuglodon atrox Andrews, 1906; D. intermedius (Dart, 1923); Prozeuglodon stromeri, Kellogg, 1928; †D. stromeri (Kellogg, 1928) Kellogg, 1936] [=?? Basilosaurus isis??] | |?- †D. zitteli (Stromer, 1903) | |-- †D. serratus Gibbes, 1845 [Zeuglodon serratum Abel 1913] | |?- †D.? sensitivus (Dart, 1923) | `?- †D.? elliotsmithii (Dart, 1923) `--+-- †Ancalecetus simonsi Gingerich & Uhen, 1996 `--+--+-- †Zygorhiza kochii (Reichenbach, 1847) True, 1908 | `-- †Pontogeneus brachyspondylus (Müller, 1849) [“Zeuglodon” brachyspondylus Müller, 1849] `--+-- †Saghacetus osiris (Dames, 1894) Gingerich, 1992 [Zeuglodon osiris Dames, 1894; Dorudon osiris (Dames, 1894)] `--+-- †Chrysocetus healyorum Uhen & Gingerich, 2001 `--o Autoceta sensu McKenna & Bell, 1997 |?- †Ocucaeja picklingi Uhen et al., 2011 `--+?-o †Kekenodontidae Mitchell, 1989 | |?- †Phococetus vasconum (Delfortrie, 1873) Gervais, 1876 | `?- †Kekenodon onamata Hector, 1881 `--o Neoceti [Cetacea sensu stricto] (modern whales; nykyvalaat) |-- Mysticeti Flower, 1864 (baleen whales; hetulavalaat) `-- Odontoceti Flower, 1867 (toothed whales; hammasvalaat)
Cetacea incertae sedis & nomina dubia
Pelagiceti Uhen, 2008
†Pontobasilus tuberculatus Leidy, 1873
†Platyosphys Brandt, 1873 [nomen dubium]
†P. paulsonii Brandt, 1873 [type lost in WWII]
†P. einori Gritsenko, 2001
†Microzeuglodon caucasicum (Lydekker, 1893) Stromer, 1903 [nomen dubium]
†Pachycetus Van Beneden, 1883 [nomen dubium]
†P. humilis Van Beneden, 1883
†P. robustus Van Beneden, 1883
†Basilosauridae Cope, 1868
After Martínez-Cáceres & de Muizon, 2011<==o Cetacea |-- †Pakicetidae `--+-- †Ambulocetidae `--+-- †Remingtonocetidae `==o “Protocetidae” |--+-- †Protocetus | `-- †Maiacetus `--+-- †Georgiacetus `--o Pelagiceti |--o Neoceti | |-- Mysticeti | `-- Odontoceti `--o †Basilosauridae |--+-- †Zygorhiza | `-- †Chrysocetus `--+--+-- †Ancalecetus | `-- †Saghacetus `--+-- †Cynthiacetus `--+-- †Dorudon `-- †Basilosaurus Reference(s):
|
After Gol'din & Zvonok, 2013<==o Cetacea |--o †Pakicetidae | `-- †Pakicetus `==o †Protocetidae |-- †Maiacetus `--+-- †Rodhocetus `--+-- †Artiocetus `--+-- †Qaisracetus `--+-- †Aegyptocetus |-- †Protocetus |-- †Carolinacetus |-- †Gaviacetus `--+-- †Eocetus |-- †Georgiacetus `--+-- †Babiacetus `--o Pelagiceti |--+-- †Ocucajea | `--+-- †Kekenodon | `-- Neoceti `--o †Basilosauridae |-- †Supayacetus `--+-- †Basilotritus `--+-- †Zygorhiza `--+-- †Chrysocetus `--+--+-- †Ancalecetus | `-- †Saghacetus `--+-- †Cynthiacetus `--+-- †Dorudon `-- †Basilosaurus |
After Geisler, Sanders & Luo, 2005<==o Cetacea |--o †Pakicetidae | `-- †Pakicetus `--+-- †Ambulocetus `--+--+-- †Dalanistes | `-- †Remingtonocetus `==o †Protocetidae |-- †Artiocetus `--+-- †Rodhocetus `--+-- †Qaisracetus `--+-- †Protocetus |-- †Carolinacetus `--+-- †Georgiacetus `--+-- †Babiacetus `--o †Basilosauridae |--+-- †Eocetus wardii | `-- †Eocetus schweinfurthi `--+-- †Dorudon `-- †Basilosaurus |
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