After Archibald, 1996, McKenna & Bell, 1997, Janis, Archibald, Cifelli, Lucas, Schaff, Schoch & Williamson, 1998, Averianov, 2000 and de Muizon & Cifelli, 2000

<==o Ungulatomorpha Archibald, 1996
   |?- †Zhelestidae genera et species indet. [UA 8699] [U. Cret. Mad.] [Averianov, Archibald & Martin, 2003]
   |?- †Kumlestes olzha Nesov, 1985 [paraphyletic Zhelestidae Nesov, 1985]
   |?- †Kumsuperus avus Nesov, 1984 [paraphyletic Zhelestidae Nesov, 1985]
   |?-o †Sorlestes Nesov, 1985 [paraphyletic Zhelestidae Nesov, 1985]
   |  |-- †S. budan Nesov, 1985
   |  `-- †S. kara Nesov, 1993
   |?- †Taslestes inobservabilis Nesov, 1982 [paraphyletic Zhelestidae Nesov, 1985]
   |-- †Aspanlestes aptap Nesov, 1985 [paraphyletic Zhelestidae Nesov, 1985]
   `--+-- †Avitotherium utahensis Cifelli, 1990d [paraphyletic Zhelestidae Nesov, 1985]
      |?-o †Gallolestes Lillegraven, 1976
      |  |-- †G. pachymandibularis Lillegraven, 1976
      |  `-- †G. agujaensis Cifelli, 1994
      |?- †Lainodon orueetxebarriai Gheerbrant & Astibia, 1994
      |?-o †Labes Sigè, 1992, in Pol, Buscalioni, Carballeira, Francés, López Martinez, Marandat, Moratella, Sanz, Sigé, Villatte, 1992
      |  |-- †L. quintanillensis Sigé, 1992, in Pol et al., 1992
      |  `-- †L. garimondi Sigé, 1992, in Pol et al., 1992
      |?- †Valentinella vitrollense Tabuce, Vianey-Liaud & Garcia, 2004
      `--+-- †Alostera saskatchewanensis Fox, 1989 [paraphyletic Zhelestidae Nesov, 1985]
         `--+--o †"Zhelestinae"
            |  |--o †Zhelestes Nesov, 1985 [paraphyletic Zhelestidae Nesov, 1985]
            |  |  |-- †Z. temirkazyk Nesov, 1985
            |  |  `-- †Z. bezelgen Nesov, 1987
            |  `--o †Parazhelestes Nesov, 1993 [paraphyletic Zhelestidae Nesov, 1985]
            |     |-- †P. robustus Nesov, 1993 [paraphyletic Zhelestidae Nesov, 1985]
            |     `-- †P. minor Nesov, Archibald & Kielan-Jaworowska, 1998 [paraphyletic Zhelestidae Nesov, 1985]
            `--+-- †Eoungulatum kudukensis Nesov et al., 1998 [??] [paraphyletic Zhelestidae]
               `--o Ungulata Linnaeus 1766 (ungulates; kaviokyntiset)
                  |?- †Kharmerungulatum vanvaleni Prasad, Verma, Sahni, Parmar & Khosla, 2007; U.Cret. SAs. (India)
                  |?-o †Hilalia Maas, Thewissen, Sen, Kazanci & Kappelman, 2001
                  |  |-- †H. sezerorum Maas, Thewissen, Sen, Kazanci & Kappelman, 2001
                  |  `--+-- †H. saribeya Maas, Thewissen, Sen, Kazanci & Kappelman, 2001 [type of Hilalia]
                  |     `--+-- †H. selanneae Maas, Thewissen, Sen, Kazanci & Kappelman, 2001
                  |        `-- †H. robusta Maas, Thewissen, Sen, Kazanci & Kappelman, 2001
                  |?- Tubulidentata (putkihampaiset, aardwarks)
                  |?- †Dinocerata (uintatheriids)
                  |-- Altungulata [Paenungulata] (horse- and elephant-like ungulates; hevos- ja norsumaiset ungulaatit)
                  `--o Eparctocyona
                     |== †Procreodi sensu McKenna & Bell, 1997 [paraphyletic assemblage of basal ungulates??]
                     `--+-- †Bulbulodentata Archibald, 1998 [Condylartha sensu McKenna & Bell, 1997, incl. †Panameriungulata de Muizon, & Cifelli, 2000 and Meridiungulata] (etelä-amerikkalaiset ungulaatit; South American ungulates)
                        `--+-- †Arctostylopida sensu Cifelli and Schaff, 1998
                           `-- "Paraxonia" (artiodactyl-like ungulates; sorkkaeläintensukuiset ungulaatit)

Note:

Taxonomy and phylogeny of basal ungulates is quite a mess. A large number of scientist have studied these animals over the years, but there is still no consensus about the relationships of main ungulate-groups.

I have here mainly followed McKenna & Bell (1997), but it's almost certain that some basal orders they recognized, such as Procreodi and Condylartha are paraphyletic. Actually all early Tertiary ungulates were once placed in Condylartha (Carroll, 1988).

One of the fun aspects when doing taxonomic research of early ungulates, are the rather funny names given by Leigh Van Valen. OK, I'm a Tolkien-fan too, but really... Did he have to...?

 

Reference(s):