After McKenna & Bell, 1997, with fossil species from “Paleobiology database”
<==o Camelidae Gray, 1821 [Camelina Gray, 1825; Camelida Haeckel, 1866] [incl. †Protolabididae Cope, 1884; †Eschatiidae Cope, 1887] (camels; kamelit) |?-o †Miotylopus Schlaikjer, 1935 [incl. Dyseotylopus Stock, 1935] | |?- †M. gibbi (Loomis, 1911) [Oxydactylus gibbiDyseotylopus migrans Stock 1935; Miotylopus bathygnathus Schlaikjer 1935] | |?- †M. leonardi (Loomis, 1911) [Gentilicamelus leonardi; Protomeryx leonardi] | `-- †M. taylori Prothero, 1996 `--+--o †Poebrodontinae (Wilson, 1974) McKenna & Bell, 1997 [†Poebrodoninae Wilson, 1974] | |--o †Poebrodon Gazin, 1955 | | |-- †P. californicus Golz, 1976 | | `-- †P. kayi Gazin, 1955 | `?- †Hidrosotherium transpecoensis Wilson, 1974 [= Hendryomeryx defordi Wilson, 1974] `--+--o †Poebrotheriinae (Cope, 1874) McKenna & Bell, 1997 [†Poebrotheriidae Cope, 1874; †Poebrotherinae (Cope) Zittel, 1893] | |--o †Poebrotherium Leidy, 1847 [†Protomeryx Leidy, 1856] | | |-- †P. wilsoni Leidy, 1847 [†Poebrotherium andersoni Troxell, 1917] (type) | | |-- †P. chadronensis Prothero, 1996 | | |-- †P. eximium Hay, 1902 | | |-- †P. franki Wilson, 1974 | | `?- †P. labratum | |-- †Paralabis cedrensis (Matthew 1901) Lull, 1921 (non McKenna, 1966) [†Pseudolabis (Paralabis) Lull, 1921; †Gentilicamelus campestris Matthew 1901, †Paralabis matthewi Lull 1921, †Paratylopus matthewi Lull 1921, †Protomeryx campester Matthew 1901, †Pseudolabis matthewi Lull 1921, †Stenomylus campester Matthew 1901] | |--o †Paratylopus (Matthew, 1904) Matthew, 1909 [†Miolabis (Paratylopus) Matthew, 1904] | | |-- †P. labiatus (Cope, 1881) | | `-- †P. primaevus (Matthew, 1904) Matthew, 1909 | `-- †Gentilicamelus sternbergi (Cope, 1879) Loomis, 1936 [†Gomphotherium Cope, 1886, non †Gomphotherium Burmeister, 1837 a genus of Proboscidea; non †Gomphotherium Filhol, 1884, a synonym of †Tetracus Aymard, 1850, a genus of Lipotyphla] `--+-- †Pseudolabis dakotensis Matthew, 1904 [Pseudolabinae Simpson, 1945] `--+-- Camelinae Zittel, 1893 [Camelidae Gray, 1821; Camelinae Zittel, 1893] [incl. †Protolabinae Zittel, 1893] (true camels; aitokamelit) `--+--o †Aepycamelinae Webb, 1965 [†Alticamelinae Simpson, 1945; †Nothokemadidae White, 1947] | |--o †Oxydactylus Peterson, 1904 | | |-- †O. benedentatus Hay, 1924 [Procamelus benedentatus] | | |-- †O. cameloides (Wortman 1898) [Gentilicamelus cameloides; Gomphotherium cameloides; Miolabis cameloides; Paratylopus cameloides] | | |-- †O. campestris Cook, 1909 | | |-- †O. lacota Matthew, 1960 | | |-- †O. longipes Peterson, 1904 | | |-- †O. longirostris Peterson, 1911 | | |-- †O. lulli Loomis, 1911 | | `-- †O. wyomingensis Loomis, 1936 | `--+-- †Australocamelus orarius Patton, 1969 | `--+--o †Aepycamelus Macdonald, 1956 [†Alticamelus Matthew, 1901 (nomen dubium); †Altomeryx Frick, 1929 (nomen nudum)] | | |-- †A. alexandrae (Davidson, 1923) [Alticamelus alexandrae; Hesperocamelus alexandrae] | | |-- †A. bradyi Macdonald, 1956 | | |-- †A. elrodi (Douglass, 1909) [Alticamelus elrodi; Procamelus elrodi] | | |-- †A. giraffinus (Matthew, 1909) Macdonald, 1956 | | |-- †A. major (Leidy, 1886) [Auchenia major; Megatylopus major; Pliauchenia major; Procamelus major] | | |-- †A. priscus (Matthew & Cook, 1909) [Alticamelus priscus] | | |-- †A. procerus (Matthew & Cook, 1909) [Alticamelus procerus] | | |-- †A. robustus (Leidy, 1858) [Alticamelus robustus; Procamelus robustus; Alticamelus madisonius Douglass, 1899; Homocamelus caninus Leidy, 1869; Procamelus madisonius Douglass, 1899] | | `-- †A. stocki Henshaw, 1942 [Alticamelus stocki] | `--+-- †Priscocamelus wilsoni M.S. Stevens, in M.S. Stevens, J.B. Stevens & Dawson, 1969 | `--+-- †Delahomeryx browni Stevens, 1969 | `--+-- †Hesperocamelus stylodon Macdonald, 1949 | `--o †Nothokemas White, 1940 | |-- †N. floridanus (Simpson 1932) [Oxydactylus floridanus; Nothokemas grandis White; 1940 Paratylopus grandis White, 1940] | |-- †N. hidalgensis Patton, 1969 | `-- †N. waldropi Frailey, 1978 `--+--o †Miolabinae Hay, 1902 | |-- †Capricamelus gettyi Whistler & Webb, 2005 | |--o †Miolabis Hay, 1899 [Miolabis Hay vide Matthew, 1898 (nomen nudum)] | | |-- †M. californicus Maxson, 1930 | | |-- †M. fissidens (Cope, 1876) [Procamelus fissidens, Protolabis fissidens] | | |-- †M. fricki Kelly, 1992 | | |-- †M. longiceps Matthew, 1909 [Protolabis longiceps] | | |-- †M. montanus (Douglass, 1899) [Protolabis montanus] | | |-- †M. princetonianus (Sinclair 1915) [Protolabis princetonianus] | | `-- †M. transmontanus (Cope, 1879) [Protolabis transmontanus] | |-- †Nothotylopus camptognathus Patton, 1969 | `--o †Paramiolabis Kelly, 1992 | |-- †P. singularis (Matthew, 1918) Honey, 1998 [Miolabis singularis Pliauchenia singularis] | |-- †P. taylori Kelly, 1992 | `-- †P. tenuis (Matthew, 1924) [Miolabis tenuis] `--+--o †Stenomylinae Matthew, gregory & Mosenthal, in Osborn, 1968 | |--o †Stenomylus Peterson, 1906 [incl. †Pegomylus Frick & Taylor, 1968] | | |-- †(Pegomylus) keelinensis Frick & Taylor, 1968 | | `--o †(Stenomylus) Peterson, 1906 | | |-- †S. (S.) crassipes Loomis, 1910 | | |-- †S. (S.) gracilis Peterson, 1906 | | |-- †S. (S.) hitchcocki Loomis, 1910 | | |-- †S. (S.) taylori Cassiliano, 2008 | | `-- †S. (S.) tubutamensis Ferrusquia-Villafranca, 1990 | `--+-- †Blickomylus galushai Frick & Taylor, 1968 | `-- †Rakomylus raki Frick, 1937 `--o †Floridatragulinae Maglio, 1966 |-- †Cuyamacamelus jamesi Kelly, 1992 `==o †Aguascalientia Stevens, 1977 [paraphyletic sensu Rincon et al., 2012] |-- †A. panamaensis Rincon, Bloch, Suarez, MacFadden & Jaramillo, 2012 `--+-- †A. minuta Rincon, Bloch, Suarez, MacFadden & Jaramillo, 2012 |-- †A. wilsoni (Dalquest & Mooser, 1974) [Miotylopus wilsoni Dalquest & Mooser, 1974] `--o †Floridatragulus White, 1940 [Hypermekops White 1942] |-- †F. dolichanthereus White, 1940 [Floridatragulus barbouri White 1947; Hypermekops olseni White 1942] |-- †F. hesperus Patton, 1969 |-- †F. nanus Patton, 1969 `-- †F. texanus Patton, 1969
Reference(s):
- Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698. - Carroll, R. L., 1988: Appendix. 594-648.
in Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698. - McKenna, M. C. & Bell, S. K., (eds.) 1997: Classification of mammals – above the species level.
–Columbia University Press, New York, 1997, xii-631 - Nowak, R. M., 1991: Walker's Mammals of the World; Part 1.
–The Johns Hopkins University Press, Baltimore and London, 1991, xlviii-642-lxiii - Nowak, R. M., 1991: Walker's mammals of the World; Part 2.
–The Johns Hopkins University Press, Baltimore and London, 1991, xii-643-1629 - Rincon, A. F., Bloch, J. I., Suarez, C., MacFadden, B. J. & Jaramillo, C. A., 2012: New Floridatragulines (Mammalia, Camelidae) from the Early Miocene Las Cascadas Formation, Panama.
–Journal of Vertebrate Paleontology: Vol. 32, #2, pp. 456-475 [doi: 10.1080/02724634.2012.635736] - Voorhies, M. R. & Corner, R. G., 1986: Megatylopus(?) cochrani (Mammalia: Camelidae): A re-evaluation.
–Journal of Vertebrate Paleontology: Vol. 6, #1, pp. 65-75