After Ahlberg & Johanson, 1998, and Long, Young, Holland, Senden & Fitzgerald, 2006
Family-level taxonomy of non-tetrapod tetrapodomorphs<==o Tetrapodomorpha Ahlberg, 1991 [Osteolepimorpha] (Terrestrial vertebrate-like sarcopterygians; maaselkärankaismaiset lihaseväiset) |-- †Tungsenia paradoxa Lu, Zhu, Long, Zhao, Senden, Jia & Qiao, 2012; E.Dev. SEAs. `--+-- †Kenichthys campbelli Chang & Zhu, 1993; E.Dev. SEAs. `--+-- †Rhizodontida Andrews & Westoll, 1970 `--o Osteolepidida Boulenger, 1901 [Osteolepiformes Berg, 1937 [non Jarvik, 1942]] |--o †Osteolepididae Cope, 1889 | |?- †Amadeodipterus kencampbelli Young & Schultze, 2005 | |?- †Luckeus abudda Young & Schultze, 2005 | |--o †Osteolepis Agassiz, 1835(?) [or Valenciennes, 1829] [Pleiopterus, Pliopterus, Triplopterus, Tripterus] M. Dev. NAs. Ant. M.-U. Dev. Eu. | | |-- †O. arenatus | | |-- †O. brevis | | |-- †O. macrolepidota (Sedgwick & Murchison, 1829)(?) [Osteolepis macrolepidotus; Thursius macrolepidotus (Sedgwick & Murchison, 1829)] | | |-- †O. major | | `-- †O. panderi | `--+-- †Muranjilepis winterensis Young & Schultze, 2005 | `--o †Thursius Sedgwick & Murchison, 1828 [non? Traquair, 1888] | |?- †T.? minor Jarvik, 1985 | |-- †T. estonicus Vorobyeva, 1977 | |-- †T. pholidotus Traquair, 1888 | |-- †T. talsiensis Traquair, 1888 | |-- †T. truncatus Traquair, 1888 | |-- †T. wudingensis Fan, 1992 | `-- †T. macrolepidotes (Sedgwick & Murchison, 1829) |?-o †Thysanolepidae [= Megalichthyidae??] | |-- †Thysanolepis micans Vorobyeva, 1977 | `-- †Vorobjevaia dolonodon Young, Long & Ritchie, 1992; L.Dev. Ant. `--+--o †Megalichthyidae Long, 1985(?) | |?- †Megistolepis klementzi (Obruchev, 1955) Vorobjeva, 1977 | |?- †Megapomus markovskyi Vorobyeva, 1977 | |-- †Mahalalepis resima Young, Long & Ritchie, 1992; L.Dev. Aust. | |?- †Sengoerichthys ottoman Janvier, Clement & Cloutier, 2007 | |-- †Ectosteorhachis nitidus Cope, 1880? | `--+-- †Cladarosymblema narrienense Fox et al., 1995 | `--o †Megalichthys | |-- †M. hibberti Agassiz, 1843 | |-- †M. laticeps Traquair, 1884 | |?- †M. agassizianus Lohest, 1889 | |?- †M. coccolepis Young, 1870 | |?- †M. intermedius Woodward, 1891 | `?- †M. macropomus Cope, 1892 `--+-- †Medoevia lata Lebedev, 1995 `--+-- †Gogonasus andrewsae Long, 1985; L.Dev. Aust. |--o †Canowindridae Young, Long & Ritchie, 1992 [Canowindididae] | |?- †Owensia chooi Holland, 2009 | |-- †Beelarongia patrichae Long, 1987 | `--+-- †Koharalepis jarviki Young, Long & Ritchie, 1992 [Koharolepis?? (lapsus calami?)] L.Dev. Ant. | `-- †Canowindra grossi Thomson, 1973 `--+--o †Gyroptychius Jarvik, 1948 | |-- †G.? australis Young & Gorter, 1981 | |-- †G. agassizi Jarvik, 1948 | |-- †G. milleri Jarvik, 1948 | |-- †G. groenlandicus Jarvik, 1950 | `-- †G. dolichotatus Jarvik, 1985 `--o Eotetrapodiformes Swartz, 2012 |-- †Tristichopteridae Cope, 1889 `--o Elpistostegalia Camp & Allison, 1961 [Panderichthyida Vorobyeva, 1981] |-- †Tinirau clackae Swartz, 2012, M.Dev. WNAm. |?- †Parapanderichthys stolbovi (Vorobyeva, 1960) Vorobyeva, 1992 [Panderichthys stolbovi Vorobyeva, 1960] `--+--o †Platycephalichthys Vorobyeva, 1959 [previously treated as tristichopterid] | |-- †P. skuenicus Vorobyeva, 1962 | |-- †P. bischoffi Vorobyeva, 1959 | `-- †P. rohoni Vorobyeva, 1962 `--+-- †Panderichthys rhombolepis Gross, 1941 [incl.? Panderichthys bystrowi Gross, 1941] [Panderichthyidae Vorobyeva, 1968] `--+-- †Elpistostege watsoni Westoll, 1938 [Elpistostegidae] |-- †Howittichthys warrenae Long & Holland 2008 |-- †Livoniana multidentata Ahlberg, Luksevic & Mark-Kurik, 2000 |-- †Tiktaalik roseae Daeschler, Shubin & Jenkins, 2006 `-- Stegocephali Laurin, 1998a [Tetrapoda Haworth, 1825 sensu Goodrich 1930, and Laurin & Reisz, 1997] (tetrapods = vertebrates with four recognizible limbs; maaselkärankaiset)
Osteolepida incertae sedis [taxa not treated by Ahlberg & Johanson, 1998]:
†Bogdanovia orientalis Obrucheva 1955; L.Dev. CAs. [has been treated as Coelacanthinimorph sarcopterygian]
†Canningius groenlandicus Säve-Söderbergh, 1937
†Chrysolepis [non Chrysolepis (chinquapins), a genus of fabaceaen plants (Fabaceae)]
†Geiserolepis
†Latvius grewingki (Gross, 1933)
†Lohsania utahensis Vaughn, 1962
†Megadonichthys kurikae Vorobyeva, 1962
†Platyethmoidia antarctica Young, Long & Ritchie, 1992; L.Dev. Ant.
†Shirolepis ananjevi Vorobeva, 1977
†Sterropterygion brandei Thomson, 1972
†Thaumatolepis edelsteini Obruchev, 1941
†Rhizodopsidae Long, 1999
†Rhizodopsis Young, 1866
†Callistiopterus clappi Romer, 1942
†Marsdenichthys longioccipitus Long, 1985; L.Dev. Aust.
†Taeniolepis trautseholdii Chabakov, 1927
After Swartz, 2012<==o Tetrapodomorpha |-- †Kenichthys `--+-- †Rhizodontida `--o Osteolepidida |--+-- †Marsdenichthys | `--o †Canowindridae[?] | |-- †Canowindra | `--+-- †Koharalepis | `-- †Beelarongia `--+--+-- †Gogonasus | `--+-- †Gyroptychius | `--+-- †Osteolepis | `--+-- †Medoevia | `-- †Megalichthyidae `--o Eotetrapodiformes |-- †Tristichopteridae `--o Elpistostegalia |-- †Tinirau `--+-- †Platycephalichthys `--+-- †Panderichthys `--+--o †Elpistostegidae[?] | |-- †Elpistostege | `-- †Tiktaalik `-- Tetrapoda Reference(s):
|
Reference(s):
- Ahlberg, P. E., Clack, J. A. & Luksevics, E. 1996: Rapib braincase evolution between Panderichthyes and the earliest tetrapods.
–Nature: vol. 381, 2 May, pp. 61-64 - Ahlberg, P. E., Luksevics, E. & Mark-Kurik, E., 2000: A near-tetrapod from the Baltic Middle Devonian.
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–W. H. Freeman and company, New York, 1988, 698 - Carroll, R. L. 1996: Revealing the patterns of macroevolution.
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–Johns Hopkins University Press, Baltimore & London, pp. 1-223 - Long, J. A., Young, G. C., Holland, T., Senden, T. J. & Fitzgerald, E. M. G., 2006: An exceptional Devonian fish from Australia sheds light on tetrapod origins.
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–Nature: Vol. 394, 6 August, pp. 569-573 - Swartz, B., 2012: A Marine Stem-Tetrapod from the Devonian of Western North America.
–Plos One: Vol. 7, #3, pp. e33683(1-11) [doi: 10.1371/journal.pone.0033683] - Young, G. C., Long, J. A. & Ritchie, A., 1992: Crossopterygian fishes from the Devonian of Antarctica: systematics, relationships and biogeographic signifcance.
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