After Stanley, Bauer, Jackman, Branch & Mouton, 2011, with Recent species from Uetz, Etzold & Chenna, 1995-2012 [“The Reptile Database”]
<==o Cordylidae Mertens, 1937 (non Gray, 1838?, non Fitzinger, 1826?) (vyöliskot) |?- †Cordylidae indet. [Augé, 2005] [Pseudolacerta sp. Augé et al., 1997] |?- †Konkasaurus mahalana Krause, Evans & Gao, 2003; U. Cret. Mad. |--o Platysaurinae Stanley, Bauer, Jackman, Branch & Mouton, 2011 | `--o Platysaurus Smith, 1844 | |?- P. guttatus Smith, 1849 (Lesser Flat Lizard) | |?- P. imperator Broadley, 1962 (Imperial Flat Lizard) | |?=o P. intermedius Matschie, 1891 (Common Flat Lizard) | | |-- P. i. inopinus Jacobsen, 1994 | | |-- P. i. natalensis Fitzsimons, 1948 | | |-- P. i. nyasae Loveridge, 1953 | | |-- P. i. parvus Broadley, 1976 | | |-- P. i. rhodesianus Fitzsimons, 1941 | | |-- P. i. subniger Broadley, 1962 | | `-- P. i. wilhelmi Hewitt, 1909 | |?- P. lebomboensis Jacobsen, 1994 (Lebombo Flat Lizard) | |?-o P. maculatus Broadley, 1965 (Spotted Flat Lizard) | | |-- P. m. maculatus Broadley, 1965 | | `-- P. m. lineicauda Broadley, 1965 | |?- P. monotropis Jacobsen, 1994 | |?- P. ocellatus Broadley, 1962 (Chimanimani Flat Lizard) | |?-o P. orientalis Fitzsimons, 1941 | | |-- P. o. orientalis Fitzsimons, 1941 | | `-- P. o. fitzsimonsi Loveridge, 1944 | |?-o P. pungweensis Broadley, 1959 (Pungwe Flat Lizard) | | |-- P. p. pungweensis Broadley, 1959 | | `-- P. p. blakei Broadley, 1964 | |?- P. relictus Broadley, 1976 (Transvaal Flat Lizard) | |?- P. torquatus Peters, 1879 (Collared Flat Lizard) | |-- P. mitchelli Loveridge, 1953 (Mitchell's Flat Lizard) | `--+--+-- P. capensis Smith, 1844 (Cape Flat Lizard) | | `-- P. broadleyi Branch & Whiting, 1997 (Broadley's Flat Lizard) | `--+-- P. intermedius nigrescens Broadley, 1981 | `--+-- P. minor Fitzsimons, 1930 | `-- P. intermedius intermedius Matschie, 1891 `--o Cordylinae Mertens, 1937 (varsinaiset vyöliskot) |--o Smaug Stanley, Bauer, Jackman, Branch & Mouton, 2011 [Cordylus in partim] | |-- S. giganteus (Smith, 1844) [Cordylus giganteus] (Giant Girdled Lizard, Sungazer; Ouvolk; jättiläisvyölisko) | `--+-- S. warreni depressus (Fitzsimons, 1930) (Zoutpansberg Girdled Lizard) | `--+--+-- S. breyeri (van Dam, 1921) (Waterberg Girdled Lizard) | | `-- S. vandami (Fitzsimons, 1930) (Van Dam's Girdled Lizard) | `--+--+-- S. warreni warreni (Boulenger, 1908) (Warren's Girdled Lizard) | | `-- S. warreni barbertonensis (van Dam, 1921) (Barberton Girdled Lizard) | `--+-- S. mossambicus (Fitzsimons, 1958) (Gorongosa Girdled Lizard) | `-- S. regius (Broadley, 1962) (Regal Girdled Lizard) `--+--+--+-- Ninurta coeruleopunctatus (Hewitt & Methuen, 1913) Stanley, Bauer, Jackman, Branch & Mouton, 2011 [Cordylus coeruleopunctatus] (Blue-spotted Girdled Lizard) | | `--o Chamaesaura Schneider, 1801 (Legless Girdled Lizards; vajaaraajaiset vyöliskot) | | |?- C. macrolepis (Cope, 1862) (Large-scale Snake Lizard) | | |?- C. miopropus Boulenger, 1894 | | |-- C. aenea (Fitzinger, 1843) (Transvaal Snake Lizard) | | `--+-- C. tenuior Günther, 1895 | | `--o C. anguina (Linnaeus, 1758) (Cape Snake Lizard) | | |-- C. a. anguina (Linnaeus, 1758) | | `-- C. a. oligopholis Laurent, 1964 | `--o Pseudocordylus Smith, 1838 | |-- P. langi Loveridge, 1944 (Lang's Crag Lizard) | `--+--+-- P. sp. “Transkei” | | `--o P. microlepidotus (Cuvier, 1829) | | |-- P. m. microlepidotus (Cuvier 1829) (Western Cape Crag Lizard) | | |-- P. m. fasciatus (Smith, 1838) (Eastern Cape Crag Lizard) | | `-- P. m. namaquensis Hewitt, 1927 | `--+--+-- P. melanotus (Smith, 1838) (Highveld Crag Lizard) | | `-- P. transvaalensis Fitzsimons, 1943 (Northern Crag Lizard) | `--+== P. subviridis (Smith, 1838) (Drakensberg Crag Lizard) | `-- P. spinosus Fitzsimons, 1947 (Spiny Crag Lizard) `--+--+-- Ouroborus cataphractus (Boie, 1828) Stanley, Bauer, Jackman, Branch & Mouton, 2011 [Cordylus cataphractus] (Armadillo Girdled Lizard; panssarivyölisko) | `--o Karusasaurus Stanley, Bauer, Jackman, Branch & Mouton, 2011 [Cordylus in partim] | |-- K. jordani (Parker, 1936) (Jordan's Girdled Lizard) | `-- K. polyzonus (Smith, 1838) (Karoo Girdled Lizard ) `--+--o Namazonurus Stanley, Bauer, Jackman, Branch & Mouton, 2011 [Cordylus in partim] | |--+-- N. lawrenci (Fitzsimons, 1939) (Lawrence's Girdled Lizard) | | `-- N. peersi (Hewitt, 1932) (Peers' Girdled Lizard) | `--+-- N. namaquensis (Methuen & Hewitt, 1914) (Namaqua Girdled Lizard) | `--+-- N. campbelli (Fitzsimons, 1938) (Campbell's Girdled Lizard) | `-- N. pustulatus (Peters, 1862) (Herero Girdled Lizard) `--+--o Hemicordylus Smith, 1838 | |-- H. capensis (Smith, 1838) [Pseudocordylus capensis] (Graceful Crag Lizard) | `-- H. nebulosus (Mouton & van Wyk, 1995) [Pseudocordylus nebulosus] (Dwarf Crag Lizard) `--o Cordylus Laurenti, 1768 [Zonurus] (vyöliskot) |?- C. angolensis (Bocage, 1895) (Angolan Girdled Lizard) |?- C. marunguensis Greenbaum, Stanley, Kusamba, Moninga, Goldberg & Bursey, 2012 (Marungu Girdled Lizard) |?- C. nyikae Broadley & Mouton, 2000 (Nyika Girdled Lizard) |?- C. rivae (Boulenger, 1896) (Ethiopian Girdled Lizard) |--+-- C. vittifer (Reichenow, 1887) (Transvaal Girdled Lizard) | `--+--+-- C. jonesii (Boulenger, 1891) (Limpopo Girdled Lizard) | | `--+-- C. machadoi Laurent, 1964 (Machado's Girdled Lizard) | | `-- C. rhodesianus (Hewitt, 1933) (Zimbabwean Girdled Lizard) | `--+-- C. beraduccii Broadley & Branch, 2002 (Maasai Girdled Lizard) | `--+-- C. ukingensis (Loveridge, 1932) (Ukinga Girdled Lizard) | `--+-- C. tropidosternum (Cope, 1869) (Tropical Girdled Lizard) | `-- C. meculae Branch, Rödel & Marais, 2005 (Mecula Girdled Lizard) `--+--+-- C. macropholis (Boulenger, 1910) (Large-scaled Girdled Lizard) | `--+-- C. mclachlani Mouton, 1986 (McLachlan's Girdled Lizard) | `--+?- C. cloetei Mouton & van Myk, 1994 (Cloete's Girdled Lizard) | |-- C. imkeae Mouton & van Myk, 1994 (Rooiberg Girdled Lizard) | `--+-- C. aridus Mouton & van Myk, 1994 (Dwarf Karoo Girdled Lizard) | `-- C. minor Fitzsimons, 1943 (Dwarf Girdled Lizard) `--+-- C. niger Cuvier, 1829 (Black Girdled Lizard) `--+-- C. oelofseni Mouton & van Myk, 1990 (Oelofsen's Girdled Lizard) `-- C. cordylus (Linnaeus, 1758) [incl. C. tasmani (Power, 1930)] (Cape Girdled Lizard)
Alternative taxonomies/phylogenies for Cordylidae
After Lang, 1991, and Krause, Evans & Gao, 2003<==o Cordylidae |?- †Cordylidae indet. [Augé, 2005] [Pseudolacerta sp. Augé et al., 1997] |?- †Konkasaurus mahalana Krause, Evans & Gao, 2003; U. Cret. Mad. |--o Chamaesaurinae | `-- Chamaesaura `--o Cordylinae |--o Cordylini | `-- Cordylus `--o Pseudocordylini |-- Platysaurus `-- Pseudocordylus |
After Frost et al., 2001 (from Stanley, Bauer, Jackman, Branch & Mouton, 2011)
<==o Cordylidae
|--o Platysaurus
| |-- P. rhodesianus
| `-- P. monotropis
`--+--+-- Cordylus cataphractus
| `--+-- Cordylus jordani
| `-- Cordylus polyzonus
`--+-- Chamaesaura aenea
`--+-- Cordylus warreni
`--+-- Cordylus giganteus
`--+--+-- Cordylus coeruleopunctatus
| `--+-- Pseudocordylus capensis
| `-- Pseudocordylus nebulosus
`--+--+-- Cordylus lawrenci
| `--+-- Cordylus vittifer
| `-- Cordylus peersi
`--+--+-- Pseudocordylus melanotus
| `--+-- Pseudocordylus microlepidotus
| `-- Pseudocordylus namaquensis
`--+-- Cordylus tropidosternum
`--+--+-- Cordylus macropholis
| `-- Cordylus minor
`--+-- Cordylus niger
`--+-- Cordylus oelofseni
`-- Cordylus cordylus
|
Reference(s):
- Augé, M., 2003: La faune de Lacertilia (Reptilia, Squamata) de l'Éocène inférieur de Prémontré (Bassin de Paris, France)
[The lizard fauna (Reptilia, Squamata) from the lower Eocene locality of Prémontré (Paris Basin, France)].
–Geodiversitas: Vol. 25, #3, pp. 539-574 - Krause, D. W., Evans, S. E. & Gao, K.-Q., 2003: First definitive record of Mesozoic lizards from Madagascar.
–Journal of Vertebrate Paleontology: Vol. 23, #4, pp. 842-856 - Stanley, E. L., Bauer, A. M., Jackman, T. R., Branch, W. R. & Mouton, P. L. F. N., 2011: Between a rock and a hard polytomy: Rapid radiation in the rupicolous girdled lizards (Squamata: Cordylidae).
–Molecular Phylogenetics and Evolution: Vol. 58, #1, pp. 53-70 [doi: 10.1016/j.ympev.2010.08.024] - Uetz, P., Etzold, T. & Chenna, R., 1995-2012: The Reptile Database [former “EMBL Reptile Database”.].
–iNet: http://www.reptile-database.org/