After Langer & Schultz, 2000, Hone & Benton, 2008, Langer, Montefeltro, Hone, Whatley & Schultz, 2010, and Ezcurra, Montefeltro & Butler, 2016
<==o †Rhynchosauria Osborn, 1903 (rhynchosaurs; rynkosaurit) |?- †Noteosuchus colletti (Watson, 1912) Broom, 1925 [Eosuchus colletti Watson, 1912] [preoccupied by Eosuchus lerichei Dollo, 1907, a gavialoid crocodylian] |?- †Colobops noviportensis Pritchard, Gauthier, Hanson, Bever & Bhullar, 2018 |-- †Mesosuchus browni Watson, 1912; E.Trias. SAf. [“Mesosuchinae”] `--+-- †Howesia browni Broom, 1905; E.Trias. SAf. `--+-- †Eohyosaurus wolvaardti Butler, Ezcurra, Montefeltro, Samathi & Sobral, 2015; M.Trias. SAf. `--o †Rhynchosauridae sensu Butler et al., 2015 [incl. Hyperdapedontidae Lydekker, 1885] |?- †Eifelosaurus triadicus Jaekel, 1904; E.Trias. Eu. |?- †Mesodapedon kuttyi Chatterjee, 1980; L.Trias. SAs. |--o †Rhynchosaurinae | `-- †Rhynchosaurus articeps Owen, 1841; M.-L.Trias. `--o †Hyperdapedontidae (Lydekker, 1885) Langer, Montefeltro, Hone, Whatley & Schultz, 2010 [Hyperdapedontinae Chatterjee, 1969] `--+-- †Langeronyx brodiei (Benton, 1990) Ezcurra, Montefeltro & Butler, 2016 [Rhynchosaurus brodiei Benton, 1990] `--+--o †Stenaulorhynchinae sensu Ezcurra, Montefeltro & Butler, 2016 | |-- †Stenaulorhynchys stockleyi (Haughton, 1932) Huene, 1938 [Hyperodepedon stockleyi; Scaphonyx stockleyi; Supradapedon stockleyi] M.Trias. EAf. | `-- “Mariante rhynchosaur” `--+-- †Bentonyx sidensis Langer, Montefeltro, Hone, Whatley & Schultz, 2010 `--+-- †Fodonyx spenceri (Benton, 1990) Hone & Benton, 2008 [Rhynchosaurus spenceri Benton, 1990] `--+-- †Ammorhynchus navajoi Nesbitt & Whatley, 2004; M.Trias. SWNAm. `--o †Hyperdapedontidae (Lydekker, 1885) Langer, Montefeltro, Hone, Whatley & Schultz, 2010 |-- †Isalorhynchus genovefae Buffetaut, 1983 [=? Hyperodepedon Huxley, 1859 (Langer et al., 2000] `--o †Hyperdapedontinae (Lydekker, 1885) Chatterjee, 1969 sensu Langer & Schultz, 2000 |-- †Teyumbaita sulcognathus (Azevedo & Schultz, 1987) Montefeltro, Langer & Schultz, 2010 [“Scaphonyx” sulcognathus Azevedo & Schultz, 1987] |-- †Otischalia elderae Hunt & Lucas, 1991 `--o †Hyperodepedon Huxley, 1859 [Stenometopon Boulenger, 1903; Scaphonyx Woodward, 1907; Macrocephalosaurus Tupi-Caldas, 1933; Paradapedon Huene, 1938; Cephalonia Huene, 1942; Supradapedon Chatterjee, 1980] L.Trias. Eu. Ind. SAs. SAm. |?- †H. australis (Huene, 1926) [Scaphonyx australis Huene, 1926] (nomen dubium?) |?- †H. lotziana (Huene, 1942) [Cephalonia lotziana Huene, 1942] (nomen dubium?) |-- †H. huenei Langer & Schultz, 2000 [Scaphonyx fischeri Woodward, 1907 in partim] L.Trias. SAm. `--+-- †H. mariensis (Tupi-Caldas, 1933) L.Trias. SAm. |--+-- †H. huxleyi Lydekker, 1881 [Paradapedon huxleyi (Lydekker, 1881) Huene, 1938], L.Trias. Ind. | `-- †H. gordoni Huxley, 1859, L.Trias. WEu. `--+?- †H. fischeri (Woodward, 1907) sensu Langer & Schultz, 2000 [Scaphonyx fischeri Woodward, 1907; Cephalonia; Cephalastron; Cephalastronius; Scaphonychimus] M. Trias. SA. [nomen dubium?] `-- †H. sanjuanensis (Sill, 1970) sensu Langer & Schultz, 2000 [Scaphonyx sanjuanensis Sill, 1970]
After Ezcurra, Montefeltro & Butler, 2016<==o †Rhynchosauria |-- †Noteosuchus colletti |-- †Mesosuchus browni |-- †Howesia browni |-- †Eohyosaurus wolvaardti `--o †Rhynchosauridae |-- †Rhynchosaurus articeps `--+--o †Stenaulorhynchinae | |-- “Mariante rhynchosaur” | `-- †Stenaulorhynchys stockleyi `--+-- †Langeronyx brodiei `--+-- †Bentonyx sidensis `--+-- †Fodonyx spenceri `--o †Hyperdapedontinae |-- †Isalorhynchus genovefae `--+-- †Teyumbaita sulcognathus `-- †Hyperodepedon Reference(s):
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After Langer, Montefeltro, Hone, Whatley & Schultz, 2010<==o †Rhynchosauria |-- †Rhynchosaurus articeps `--o †Hyperdapedontidae |--+-- “Mariante rhynchosaur” | `-- †Stenaulorhynchys stockleyi |-- †Bentonyx sidensis `--+-- †Fodonyx spenceri `--o †Hyperdapedontinae |-- †Isalorhynchus genovefae `--+-- †Teyumbaita sulcognathus `-- †Hyperodepedon Reference(s):
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Reference(s):
- Butler, R. J., Ezcurra, M. D., Montefeltro, F. C., Samathi, A. & Sobral, G., 2015: A new species of basal rhynchosaur (Diapsida: Archosauromorpha) from the early Middle Triassic of South Africa, and the early evolution of Rhynchosauria.
–Zoological Journal of the Linnean Society: Vol. 174, #3, pp. 571-588 [doi: 10.1111/zoj.12246] - Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698. - Carroll, R. L., 1988: Appendix. 594-648.
in Carroll, R. L. 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698 - Ezcurra, M. D., Montefeltro, F. & Butler, R. J., 2016: The early evolution of rhynchosaurs.
–Frontiers in Ecology and Evolution: Vol. 3, #142 [doi: 10.3389/fevo.2015.00142] - Ezcurra, M. D., Trotteyn, M. J., Fiorelli, L. E., Baczko, M. B., Taborda, J. R. A., Iberlucea, M. & Desojo, J. B., 2013: The oldest rhynchosaur from Argentina: a Middle Triassic rhynchosaurid from the Chañares Formation (Ischigualasto–Villa Unión Basin, La Rioja Province).
–Paläontologische Zeitschrift: in press pp. 1-8 [doi: 10.1007/s12542-013-0203-3] - Hone, D. W. E. & Benton, M. J., 2008: A new genus of rhynchosaur from the Middle Triassic of south-west England.
–Palaeontology: Vol. 51, #1, pp. 95-115 [doi: 10.1111/j.1475-4983.2007.00739.x] - Langer, M. C., Montefeltro, F. C., Hone, D. E., Whatley, R. & Schultz, C. L., 2010: On Fodonyx spenceri and a New Rhynchosaur from the Middle Triassic of Devon.
–Journal of Vertebrate Paleontology: Vol. 30, #6, pp. 1884-1888 [doi: 10.1080/02724634.2010.521901] - Langer, M. C. & Schultz, C. L., 2000: A new species of the Late Triassic rhynchosaur Hyperodapedon from the Santa maria Formation of South Brazil.
–Paleontology: Vol. 43, #4, pp. 633-652 - Montefeltro, F. C., Langer, M. C. & Schultz, C. L., 2010: Cranial anatomy of a new genus of hyperodapedontine rhynchosaur (Diapsida, Archosauromorpha) from the Upper Triassic of southern Brazil.
–Earth and Environmental Science Transactions of the Royal Society of Edinburgh: Vol. 101, #01, pp. 27-52 [doi: doi:10.1017/S1755691010009060] - Nesbitt, S. J. & Whatley, R. L., 2004: The first discovery of a rhynchosaur from the upper Moenkopi Formation (Middle Triassic) of northern Arizona.
–PaleoBios: Vol. 24, #3, pp. 1-10 - Pritchard, A. C., Gauthier, J. A., Hanson, M., Bever, G. S. & Bhullar, B.-A. S., 2018: A tiny Triassic saurian from Connecticut and the early evolution of the diapsid feeding apparatus.
–Nature Communications: Vol. 9, #1, Art.nr. 1213, pp. (1-10) [doi: 10.1038/s41467-018-03508-1]