After Paul, 1988

<==o Theropoda
   `==o Paleotheropoda Paul, 1988 (paleotheropods) [paraphyletic]
      `==o Ceratosauria Marsh, 1884 (ceratosaurs; keratosaurit) [paraphyletic]
         |--+-- †Procompsognathidae Huene, 1929
         |  `--o †Coelophysidae Welles, 1984 sensu Paul, 1988
         |     |-- †Coelophysinae Welles, 1984 sensu Paul, 1988
         |     `--+-- †Halticosaurinae Huene, 1948 sensu Paul, 1988
         |        `-- †Spinosauridae Stromer, 1915 [incl. Baryonychidae Charig & Milner, 1986]
         `--+-- †Ceratosauridae Marsh, 1884
            `==o Intertheropoda Paul, 1988 [paraphyletic]
               |--o †Megalosauridae Huxley, 1869 [Torvosauridae]
               |  |-- †Megalosaurinae Huxley, 1869 sensu Paul, 1998
               |  `--+-- †Abelisaurinae Bonaparte & Novas, 1986 sensu Paul, 1998
               |     `-- †Noasaurinae Bonaparte & Powell, 1980 sensu Paul, 1998
               `--o †Eustreptospondylidae Paul, 1988 [according to Paul's cladogram is a paraphyletic family]
                  |-- †Eustreptospondylinae Paul, 1988
                  `--+-- †Metriacanthosaurinae Paul, 1988
                     |?- †Segisauridae Camp, 1939
                     |?- †Iliosuchidae Paul, 1988
                     `--o AVETHEROPODA Paul, 1988 [Neotheropoda Sereno]
                        |?- †Dryptosauridae Marsh, 1890
                        |--o †COMPSOGNATHIA Paul, 1988
                        |  `-- †Compsognathidae Cope, 1875 (gompsognathids; siroleuat)
                        |--+--o †COELURIA Paul, 1988
                        |  |  `-- †Coeluridae Marsh, 1881
                        |  `--o †Allosauria Paul, 1988 [Carnosauria]
                        |     `==o †Allosauridae Marsh, 1878
                        |        |-- †Ornitholestinae Paul, 1988 (ornitholestines; linnunnappaajat)
                        |        `--+==Allosaurinae Marsh, 1878 (true allosaurs; allosaurit)
                        |           `--o †Tyrannosauridae Osborn, 1906 (Tyrannisaurit)
                        |              |-- †Aublysodontinae Nopsca, 1928 [has later been noticed to be only juvelines of larger tyrannosaurs]
                        |              `-- †Tyrannosaurinae Osborn, 1906 (suurtyrannisaurit)
                        `--o PROTOAVIA Paul, 1988 [Maniraptora Gauthier, 1986]
                           `==o Archaeopterygidae Huxley, 1872 sensu Paul, 1988 [according to Paul's cladogram is a paraphyletic family]
                              |-- †Archaeopteryginae Huxley, 1872 sensu Paul, 1988 (archaeopterygines; archaeopterygit)
                              `--+-- †Dromaeosaurinae Matthew & Brown, 1922 sensu Paul, 1988 (raptors; raptorit, sirppikynnet)
                                 `--+--o †Caenagnathidae Sternberg, 1940 sensu Paul, 1988 [incl. Elmisauridae]
                                    |  |-- †Caenagnathinae Sternberg, 1940 sensu Paul, 1988
                                    |  `-- †Oviraptorinae Barsbold, 1976 sensu Paul, 1988 (oviraptors)
                                    `--+--o †Ornithomimidae Marsh, 1890 (ornithomimids; strutsisaurit)
                                       |  |-- †Garudimiminae Barsbold, 1981 sensu Paul, 1988
                                       |  |-- †Deinocheirinae Osmólska & Roniewicz, 1969 sensu Paul, 1988
                                       |  `-- †Ornithomiminae Marsh, 1890 sensu Paul, 1988 (strutsisaurit)
                                       |-- †Troodontidae Gilmore, 1924 (valesirppikynnet)
                                       `--+?- †Avimimidae Kurzanov, 1981 (avimimukset)
                                          `-- AVES Linnaeus, 1758 [Avialae Gauthier, 1986]

Note: Gregory S. Paul's wonderful book, "Predatory Dinosaurs of the World" from 1988, is a real tome of detailed anatomical information. It also includes some very beautiful artwork. Among people who draw dinosaurs, there is no greater acclaim than being called "Paulish".

Despite all this, his views of the phylogeny and evolution of theropods are highly criticized and controversial. One of his main thoughts is that Protoavians are actually secondarily flightess descendants of Archaeopteryx. One should also remember that a number of rather interresting theropods, such as Alvarezsauridae, were not known at 1988.

More recently Paul has used cladograms from Holtz. (Paul, 2002)

 

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