Mikko's Phylogeny Archive Non-Avian Archosaur Groups   †Pachycephalosauria - thick-headed dinosaurs

After Sullivan, 2003

<==o †Pachycephalosauria (Sternberg, 1945) Marya´nska & Osmólska, 1974 (thick-headed dinosaurs; paksupäät)
   |?- †Dracorex hogwartsia
   |-- †Stenopelix valdensis Meyer, 1857
   `--+-- †Wannanosaurus yansiensis
      `--o †Goyocephalia
         |-- †Goyocephale lattimorei Perle, Marya´nska & Osmólska, 1982
         `--o †Homalocephaloidea Dong, 1974 sensu Sereno, 1990 [Homalocephalidae Dong, 1974 sensu lato]
            |-- †Homalocephale calathocercos Marya´nska & Osmólska, 1974
            |-- †Ornatotholus browni (Wall & Galton, 1979) Galton & Sues, 1983 [Stegoceras browni Wall & Galton, 1979]
            `--+-- †Yaverlandia bitholus Galton, 1971c
               `--o †Pachycephalosauridae Sternberg, 1945 [Tholocephalidae]
                  |-- †Stegoceras validum (Lambe, 1902) emend. Sues & Galton, 1987 [taxonomic history]
                  `--+-- Pachycephalosauridae genera et species indet. [North American dwarf species]
                     `--o †Pachycephalosaurinae Sternberg, 1945 sensu Sereno, 2000
                        |--o "Tylocephalini"
                        |  |-- †Tylocephale gilmorei Marya´nska & Osmólska, 1974
                        |  `--o †Prenocephale Marya´nska & Osmólska, 1974 [taxonomic history]
                        |     |-- †P. prenes Marya´nska & Osmólska, 1974 [taxonomic history]
                        |     `--+-- †P. goodwini (Williamson & Carr, 2003) Sullivan, 2003 [taxonomic history]
                        |        `--+-- †P. edmontonensis (Brown & Schlaikjer, 1943) Sullivan, 2000 [taxonomic history]
                        |           `-- †P. brevis (Lambe, 1918b) Sullivan, 2000 [taxonomic history]
                        `--+--+-- †Colepiocephale lambei (Sternberg, 1945) Sullivan, 2003 [taxonomic history]
                           |  `-- †Hanssuessia sternbergi (Brown & Schlaikjer, 1943) Sullivan, 2003 [taxonomic history]
                           `--o †Pachycephalosaurini (Sternberg, 1945) Sullivan, 2003
                              |-- †Stygimoloch spinifer Galton & Sues, 1983 [Stenotholus kohleri Giffin, Gabriel & Johnson, 1987]
                              `--o †Pachycephalosaurus Brown & Schlaikjer, 1943 [Tylosteus Leidy, 1872]
                                 |?- †P. sp.
                                 `-- †P. wyomingensis (Gilmore, 1931) Brown & Schlaikjer, 1943 [Troodon wyomingensis Gilmore, 1931, incl. Tylosteus ornatus Leidy, 1872; Pachycephalosaurus grangeri Brown & Schlaikjer, 1943; Pachycephalosaurus reinheimeri Brown & Schlaikjer, 1943]


Pachycephalosauria incertae sedis:

    †Micropachycephalosaurus hongtuyanensis Dong, 1978
    †"Troodon" bexelli Bohlin, 1953
    †Gravitholus albertae Wall, & Galton, 1979 (nomen dubium)

Reference(s):

Dodson, P., 1990: Marginocephalia. 562-563.
in Weishampel, D. B., Dodson, P., & Osmólska, H. (eds.), 1990: The Dinosauria.
–University of California Press, Berkley, Los Angeles, Oxford, 1990 xvi-733

Goodwin, M. B., Buchholtz, E. A. & Johnson, R. E., 1998: Cranial anatomy and diagnosis of Stygimoloch spinifer (Ornithischia: Pachycephalosauria) with comments on cranial display structures in agonistic behavior.
–Journal of Vertebrate Paleontology: Vol. 18, #2, pp. 363-375

Keesey, T. M., 1996-2000: The Dinosauricon

–iNet: The Dinosauricon: http://dinosauricon.com/

Marya´nska, T., 1990: Pachycephalosauria. 564-577
in Weishampel, D. B., Dodson, P., & Osmólska, H. (eds.), 1990: The Dinosauria.
–University of California Press, Berkley, Los Angeles, Oxford, 1990 xvi-733

Sereno, P. C., 2000: The fossil record, systematics and evolution of pachycephalosaurs and ceratopsians from Asia. 480-516
in Benton, M. J., Shishkin, M. A., Unvin, D. M. & Kurochkin, E. N., (eds.) 2000: The Age of Dinosaurs in Russia and Mongolia.
–Cambridge University Press, Cambridge, xxxix-696

Sullivan, R. M., 2000: Stegoceras revisited.
–Journal of Vertebrate Paleontology: Vol. 20, #3, suppl. to #3, 72A

Sullivan, R. M., 2003: Revision of the dinosaur Stegoceras Lambe (Ornithischia, Pachycephalosauria).
–Journal of Vertebrate Paleontology: Vol. 23, #1, pp. 181-207

Williamson, T. E. & Carr, T. D., 2003: A new genus of derived pachycephalosaurian from western North America.
–Journal of Vertebrate Paleontology: Vol. 22, #4, pp. 779-801