After Xing, Wang, Han, Sullivan, Ma, He, Hone, Yan, Du & Xu, 2014
<==o †Hadrosauroidea (Cope, 1869b) Norman, 2002 (duck-billed dinosaurs and relatives; ankannokkasauriensukuiset hierrinhampaiset) |-- †Equijibus normani You et al., 2003a `--+-- †Xuwulong yueluni You, Li & Liu, 2011 `--+-- †Probactrosaurus gobiensis Rozhdestvensky, 1966 [incl. P. alashanicus Rozhdestvensky, 1966] [Probactrosauridae] `--+-- †Jintasaurus meniscus You & Li, 2009 |--+-- †Protohadros byrdi Head, 1998 | `-- †Eolambia caroljonesa Kirkland, 1998 `--+-- †Levnesovia transoxiana Sues & Averianov, 2009; L.Cret. CAs. `--+-- †Tanius sinensis Wiman, 1929 [incl. T. chingkankouensis Young, 1958a] `--+-- †Bactrosaurus johnsoni Gilmore, 1933a `--+-- †Gilmoreosaurus mongoliensis (Gilmore, 1933a) Brett-Surman, 1979 [Mandschurosaurus mongoliensis Gilmore, 1933a] `--+-- †Shuangmiaosaurus gilmorei You, Ji, Li & Li, 2003 [has been treated as non-Hadrosauroid Iguanodontoid] `--+-- †Nanningosaurus dashiensis Mo, Zhao, Wang, & Xu, 2007 `--+--+-- †Telmatosaurus transsylvanicus (Nopcsa, 1900) Nopcsa, 1903 [Limnosaurus transsylvanicus Nopsca, 1900; Hecatosaurus Brown, 1910] | `-- †Tethyhadros insularis Dalla Vecchia, 2009 `--+-- †Claosaurus agilis (Marsh, 1872) [Hadrosaurus agilis Marsh, 1872; Claosaurus Marsh, 1890a] `--+-- †Zhanghenglong yangchengensis Xing, Wang, Han, Sullivan, Ma, He, Hone, Yan, Du & Xu, 2014; L.Cret. EAs. |-- †Nanyangosaurus zhugeii Xu, Zhao, Huang, Li & Dong, 2000 `--o †Hadrosauridae Cope, 1869b (duck-billed dinosaurs; ankannokkasaurit) |-- †Hadrosaurinae Cope, 1869 (Lambe, 1918a) [Saurolophinae] (tavalliset ankankokkasaurit) `-- †Lambeosaurinae Parks, 1923 [Hypacrosaurinae] (helttakalloiset ankannokkasaurit)
Untreated Hadrosauroids:
†Siamodon nimngami Buffetaut & Suteethorn, 2011; E.Cret. SEAs.
†Penelopognathus weishampeli Godefroit, Li & Shang, 2005
†Altirhinus kurzanovi Norman, 1998 [incl. “Iguanodon” orientalis Rozhdestvensky, 1952] (aasiantikarisormi)
†Jeyawati rugoculus Wolfe & Kirkland, 2010
†Huehuecanauhtlus tiquichensis Ramírez-Velasco, Benammi, Prieto-Márquez, Ortega & Hernández-Rivera, 2012
†Koshisaurus katsuyama Shibata & Azuma, 2015
Hadrosauroidea incertae sedis:
†“Stegosaurus” madagascariensis
Nomina dubia & invalida
†Cionodon? kysylkumense Riabinin, 1931 (nomen dubium)
†Gilmoreosaurus arkhangelskyi Nessov, 1995 (nomen dubium)
†Hadrosaurus foulkii Leidy, 1856 (nomen dubium, Prieto-Marquez, Weishampel & Horner, 2006) [Hadrosaurinae indet.]
†Hadrosaurus tripos Cope, 1869 (nomen dubium, Baird & Horner, 1979) [a Pliocene whale bone (Belaenopteridae)]
†Hadrosaurus minor Marsh, 1870 (nomen dubium, Prieto-Marquez, Weishampel & Horner, 2006) [Hadrosauridae indet.]
†Hadrosaurus cavatus Cope, 1871 (nomen dubium, Weishampel & Horner, 1990) [Hadrosauridae indet.]
†Hadrosaurus breviceps Marsh, 1889 (nomen dubium, Prieto-Marquez, Weishampel & Horner, 2006) [Hadrosauridae indet.]
= Kritosaurus breviceps (Marsh); Lull & Wright, 1942
†Hadrosaurus paucidens Marsh, 1889 (nomen dubium, Prieto-Marquez, Weishampel & Horner, 2006) [Lambeosaurinae indet.]
†Ornithotarsus immanis Cope, 1869c (nomen dubium, Prieto-Marquez, Weishampel & Horner, 2006) [Hadrosauridae indet.]
†“Trachodon” cantabrigiensis (nomen nudum?, teeth)
†“Gilmoreosaurus” atavus (nomen nudum?, teeth)
Alternative phylogenies of basal hadrosauroids:
After Xing, Wang, Han, Sullivan, Ma, He, Hone, Yan, Du & Xu, 2014<==+-- †Jinzhousaurus `--o Hadrosauroidea |-- †Equijibus `--+-- †Xuwulong `--+-- †Probactrosaurus `--+-- †Jintasaurus |--+-- †Protohadros | `-- †Eolambia `--+-- †Levnesovia `--+-- †Tanius `--+-- †Bactrosaurus `--+-- †Gilmoreosaurus `--+-- †Shuangmiaosaurus `--+-- †Nanningosaurus `--+--+-- †Telmatosaurus | `-- †Tethyhadros `--+-- †Claosaurus `--+-- †Zhanghenglong |-- †Nanyangosaurus `--o †Hadrosauridae |-- †Hadrosaurinae `-- †Lambeosaurinae Reference(s):
|
After Prieto-Márquez, 2010<==o Hadrosauroidea |-- †Altirhinus |-- †Equijibus `--+-- †Probactrosaurus `--+--+-- †Eolambia | `-- †Protohadros `--+-- †Tanius `--+-- †Bactrosaurus `--+-- †Gilmoreosaurus `--+-- †Telmatosaurus `--+-- †Lophorhothon `--o †Hadrosauridae |-- †Hadrosaurus foulkii `--o †Saurolophidae |-- †Saurolophinae `-- †Lambeosaurinae Reference(s):
|
|
After McDonald, Kirkland, DeBlieux, Madsen, Cavin, Milner & Panzarin, 2010<==o Hadrosauroidea |-- †Altirhinus |-- †Equijibus |--+-- †Jinzhousaurus | `-- †Penelopognathus `--+-- †Eolambia |-- †Probactrosaurus `--+-- †Jeyawati `--+-- †Protohadros `--+-- †Levnesovia |-- †Nanyangosaurus |-- †Tethyhadros `--+-- †Bactrosaurus `--+-- †Gilmoreosaurus `--+-- †Tanius `--+-- †Lophorhothon `--+-- †Telmatosaurus |-- †Claosaurus `-- †Hadrosauridae s.s. Reference(s):
|
After Sues & Averianov, 2009
<==+-- †Equijibus
|-- †Probactrosaurus
`--+-- †Protohadros
`--o †Hadrosauroidea
|--+-- †Bactrosaurus
| `-- †Levnesovia
`--+-- †Tanius
`--+-- †Telmatosaurus
`--+-- †Aralosaurus
`-- †Hadrosauridae
Reference(s):
|
After Norman, 2002<==o †Hadrosauroidea |-- †Altirhinus `--+-- †Eolambia `--+-- †Protohadros `--+-- †Probactrosaurus `--+-- †Telmatosaurus `--+-- †Bactrosaurus `-- †Hadrosauridae Reference(s):
|
Reference(s):
- Buffetaut, E. & Suteethorn, V., 2011: A new iguanodontian dinosaur from the Khok Kruat Formation (Early Cretaceous, Aptian) of northeastern Thailand.
–Annales de Paléontologie: Vol. 97, #1–2, pp. 51-62 [doi: http://dx.doi.org/10.1016/j.annpal.2011.08.001] - Carpenter, K., Dilkes, D. W. & Weishampel, D. B., 1995: The dinosaurs of the Niobrara Chalk Formation (Upper Cretaceous, Kansas).
–Journal of Vertebrate Paleontology: Vol. 15, #2, pp. 275-297 - Godefroit, P., Li, H. & Shang, C.-Y., 2005: A new primitive hadrosauroid dinosaur from the early Cretaceous of Inner Mongolia (P.R. China).
–Comptes Rendus de l'Académie des Sciencies, Paris: Palevol: Vol. 4, pp. 697-705 [doi: 10.1016/j.crpv.2005.07.004] - Head, J. J., 1998: A new species of basal hadrosaurid (Dinosauria, Ornithischia) from the Cenomanian of Texas.
–Journal of Vertebrate Paleontology: Vol. 18, #4, pp. 718-738 - Keesey, T. M., 1996-2000: Hadrosauroidea – Duck-billed dinosaurs
–iNet: The Dinosauricon: http://dinosauricon.com/taxa/Hadrosauroidea.html - McDonald, A. T., Kirkland, J. I., DeBlieux, D. D., Madsen, S. K., Cavin, J., Milner, A. R. C. & Panzarin, L., 2010: New Basal Iguanodonts from the Cedar Mountain Formation of Utah and the Evolution of Thumb-Spiked Dinosaurs.
–Plos One: Vol. 5, #11, pp. e14075 (1-35) [doi: 10.1371/journal.pone.0014075] - Norman, D. B., 2002: On Asian ornithopods (Dinosauria: Ornithischia). 4. Probactrosaurus Rozhdestvensky, 1966.
–Zoological Journal of the Linnean Society: Vol. 136, #1, pp. 113-144 [doi: 10.1006/zjls.1997.0122] - Norman, D. B., & Weishampel, D. B., 1990: Iguanodontidae and related ornithopods. 510-533.
in Weishampel, D. B., Dodson, P., & Osmólska, H. (eds.), 1990: The Dinosauria.
–University of California Press, Berkley, Los Angeles, Oxford, 1990, xvi-733 - Prieto-Marquez, A., Weishampel, D. B. & Horner, J. R., 2006: The dinosaur Hadrosaurus foulkii, from the Campanian of the East Coast of North America, with a reevaluation of the genus.
–Acta Palaeontologica Polonica: Vol. 51, #1, pp. 77-98 - Ramírez-Velasco, A. A., Benammi, M., Prieto-Márquez, A., Ortega, J. A. & Hernández-Rivera, R., 2012: Huehuecanauhtlus tiquichensis, a new hadrosauroid dinosaur (Ornithischia: Ornithopoda) from the Santonian (Late Cretaceous) of Michoacán, Mexico.
–Canadian Journal of Earth Sciences: Vol. 49, #2, pp. 379-395 [doi: 10.1139/e11-062] - Shibata, M. & Azuma, Y., 2015: New basal hadrosauroid (Dinosauria: Ornithopoda) from the Lower Cretaceous Kitadani Formation, Fukui, central Japan.
–Zootaxa: Vol. 3914, #4, pp. 20 [doi: 10.11646/zootaxa.3914.4.3] - Sues, H.-D. & Averianov, A. O., 2009: A new basal hadrosauroid dinosaur from the Late Cretaceous of Uzbekistan and the early radiation of duck-billed dinosaurs.
–Proceedings of the Royal Society of London: Biological series: Vol. 276, pp. 2549-2555 [doi: 10.1098/rspb.2009.0229] - Sues, H.-D., & Norman, D. B., 1990: Hypsilophodontidae, Tenontosaurus, Dryosauridae. 498-509
in Weishampel, D. B., Dodson, P., & Osmólska, H. (eds.), 1990: The Dinosauria.
–University of California Press, Berkley, Los Angeles, Oxford, 1990, xvi-733 - Weishampel, D. B., & Norman, D. B., 1990: Hadrosauridae. 534-561.
in Weishampel, D. B., Dodson, P., & Osmólska, H. (eds.), 1990: The Dinosauria.
–University of California Press, Berkley, Los Angeles, Oxford, 1990, xvi-733 - Xing, H., Wang, D., Han, F., Sullivan, C., Ma, Q., He, Y., Hone, D. W. E., Yan, R., Du, F. & Xu, X., 2014: A New Basal Hadrosauroid Dinosaur (Dinosauria: Ornithopoda) with Transitional Features from the Late Cretaceous of Henan Province, China.
–Plos One: Vol. 9, #6, pp. e98821 [doi: 10.1371/journal.pone.0098821] - You, H.-L. & Li, D.-Q., 2009: A new basal hadrosauriform dinosaur (Ornithischia: Iguanodontia) from the Early Cretaceous of northwestern China.
–Canadian Journal of Earth Sciences: Vol. 46, #12, pp. 949-957 [doi: 10.1139/E09-067]