After Sereno, 2000, Makovicky, 2001, Xu et al., 2002, Xu et al., 2006, and Makovicky & Norell, 2006
Alternative phylogeny of Ceratopsia after Sereno, 2000, and You & Dodson, 2003
Alternative phylogeny of Ceratopsia after Makovicky, 2001
Alternative phylogeny of Ceratopsia after You & Dodson, 2003
Alternative phylogeny of Ceratopsia after Chinnery, 2004
<==o †Ceratopsia Marsh, 1890 (sarvinaamat; horned dinosaurs) |-- †Yinlong downsi Xu, Forster, Clark & Mo, 2006 `--+--o †Chaoyangsauridae Zhao, Cheng, Xu & Makovicky, 2006 | |-- †Chaoyangsaurus youngi Zhao, Cheng & Xu, 1999 | `-- †Xuanhuaceratops niei Zhao, Cheng, Xu & Makovicky, 2006 `--+-- †Psittacosauria Osborn, 1923 `--o †Neoceratopsia Sereno, 1991 [Neoceratopia] |?- †Craspedodon lonzeensis Dollo, 1883 sensu Godefroit & Lambert, 2007 [tooth-taxa, could as well be an ornithopod] |-- †Liaoceratops yanzigouensis Xu, Makovicky, Wang, Norell & You, 2002 `--+-- †Yamaceratops dorngobiensis Makovicky & Norell, 2006 `--+-- †Archaeoceratops oshimai Dong & Azuma, 1997 [“Archaeoceratopidae”] `--+--o †Leptoceratopsidae Nopcsa, 1923 | |?- †Cerasinops hodgskissi Chinnery & Horner, 2007 | |-- †Asiaceratops salsopaludalis Nesov, Kaznyshkina & Cherepanov, 1989 [nomen dubium] | `--+-- †Montanoceratops cerorhynchus (Brown & Schlaikjer, 1942) Sternberg, 1951 [Leptoceratops cerorhynchus Brown & Schlaikjer, 1942] | `--+-- †Prenoceratops pieganensis Chinnery, 2004 | `--+-- †Udanoceratops tschizhovi Kurzanov, 1992 | `-- †Leptoceratops gracilis Brown, 1914c `--o †Coronosauria Sereno, 1986 |--o †Protoceratopidae Granger & Gregory, 1923 [incl.? Bagaceratopidae Alifanov, 2003] | |?- †Ajkaceratops kozmai Ösi, Butler & Weishampel, 2010 | |-- †Gobiceratops minutus Alifanov, 2008 | |-- †Graciliceratops mongoliensis Sereno, 2000 | |-- †Bagaceratops rozhdestvenskyi Marya´nska & Osmólska, 1975 [Breviceratops kozlowskii (Marya´nska & Osmólska, 1975)] | |-- †Lamaceratops tereschenkoi Alifanov, 2003 | |-- †Magnirostris dodsoni You & Dong, 2003 | |-- †Platyceratops tatarinovi Alifanov, 2003 | `--o †Protoceratops Granger & Gregory, 1923 | |-- †P. andrewsi Granger & Gregory, 1923 | `-- †P. hellenikorhinus Lambert, Godefroit, Li, Shang & Dong, 2001 `--o †Ceratopsoidea Hay, 1902 [Ceratopoidea] `--o †Ceratopsomorpha |-- †Zuniceratops christopheri Wolfe & Kirkland, 1998 `--+-- †Turanoceratops tardabilis Nesov, Kaznyshkina & Cherepanov, 1989 `-- †Ceratopsidae Marsh, 1890a [Ceratopidae Marsh, 1890a emend. ??, Agathaumidae Cope]
Ceratopsia incertae sedis:
†Auroraceratops rugosus You, Li, Ji, Lamanna & Dodson, 2005
†Kulceratops kulensis
†Microceratops Bohlin, 1953 sensu Sereno, 2000 [Nomia dubia]
†M. gobiensis Bohlin, 1953 [Nomia dubia]
†M. sulcidens Bohlin, 1953 [Nomia dubia]
†Serendipaceratops arthurcclarkei Rich & Vickers-Rich, 2003 [An Australian leptoceratopsid?]
After Sereno, 2000, and You & Dodson, 2003<==o †Ceratopsia |-- †Psittacosauria `--o †Neoceratopsia |-- †Chaoyangsaurus `--+-- †Liaoceratops `--+-- †Archaeoceratops `--+--o †“Leptoceratopidae” | |?- †Udanoceratops | `-- †Leptoceratops `--o †Coronosauria |--o †Protoceratopidae | |-- †Graciliceratops | |-- †Bagaceratops | `-- †Protoceratops `--o †Ceratopsoidea |-- †Montanoceratops `--o †Ceratopsomorpha |-- †Zuniceratops `--+-- †Turanoceratops `-- †Ceratopsidae |
After Makovicky, 2001<==o †Ceratopsia |--+-- Ceratopsia genera et species indet. [China] [Makovicky, 2001] | `-- †Chaoyangsaurus `--+-- †Psittacosauria `--o †Neoceratopsia |-- †Archaeoceratops `--+--o †“Leptoceratopidae” | |-- †Leptoceratops | |-- †Montanoceratops | `-- †Udanoceratops `--o †Coronosauria |--o †Protoceratopidae | |?- †Graciliceratops | |-- †Bagaceratops | `-- †Protoceratops `--o †Ceratopsoidea `--o †Ceratopsomorpha |-- †Zuniceratops `--+?- †Turanoceratops `-- †Ceratopsidae |
After You & Dodson, 2003<==o †Ceratopsia |-- †Psittacosauria `--o †Neoceratopsia |-- †Chaoyangsaurus `--+-- †Liaoceratops `--+-- †Archaeoceratops `--o †Coronosauria |--o †Protoceratopidae | |-- †Graciliceratops | |-- †Bagaceratops | `-- †Protoceratops `--o †Ceratopsoidea |--o †Leptoceratopsidae | |?- †Udanoceratops | |-- †Montanoceratops | `-- †Leptoceratops `--o †Ceratopsomorpha |-- †Zuniceratops `--+-- †Turanoceratops `-- †Ceratopsidae |
After Chinnery, 2004a, b<==o †Ceratopsia |-- †Psittacosauria `--o †Neoceratopsia |--o †“Chaoyangsauridae” | |-- †Chaoyangsaurus | `-- †Asiaceratops `--+-- †Liaoceratops `--+-- †Archaeoceratops `--+--o †“Leptoceratopsidae” | |-- †Montanoceratops | `--+-- †Prenoceratops | `--+-- †Udanoceratops | `-- †Leptoceratops `--o †Coronosauria |--o †Protoceratopidae | |-- †Graciliceratops | |-- †Bagaceratops | `-- †Protoceratops `--o †Ceratopsoidea `--o †Ceratopsomorpha |-- †Zuniceratops `--+-- †Turanoceratops `-- †Ceratopsidae |
Reference(s):
- Chinnery, B. J., 2004: Description of Prenoceratops pieganensis gen. et sp. nov. (Dinosauria: Neoceratopsia) from the Two Medicine Formation of Montana.
–Journal of Vertebrate Paleontology: Vol. 24, #3, pp. 572-590 - Chinnery, B. J. & Weishampel, D. B., 1998: Montanoceratops cerorhynchus (Dinosauria: Ceratopsia) and relationships among basal neoceratopsians.
–Journal of Vertebrate Paleontology: Vol. 18, #3, pp. 369-585 - Dopson, P., 1996: The Horned Dinosaurs.
–Princeton University Press, Pinceton, New Jersey, pp. xiv-346 - Dopson, P., & Currie, P. J., 1990: Neoceratopsia. 593-618.
in Weishampel, D. B., Dodson, P., & Osmólska, H. (eds.), 1990: The Dinosauria.
–University of California Press, Berkley, Los Angeles, Oxford, 1990 xvi-733 - Godefroit, P. & Lambert, O., 2007: A re-appraisal of Craspedodon lonzeensis Dollo, 1883 from the Upper Cretaceous of Belgium: the first record of a neoceratopsian dinosaur in Europe.
–Bulletin de l'Institut Royal des Sciences Naturelles de Belgique - Sciences de la Terra / Bulletin van het Koninkljik Belgisch Instituut voor Natuurwetenschappen - Aardwetenschappen: Vol. 77, pp. 83-93 - Keesey, T. M., 1996-2000: The Dinosauricon
–iNet: http://dinosauricon.com/ - Lambert, O, Godefroit, P, Li, H, Shang, C.-Y. & Dong,Z.-M., 2001: A new species of Protoceratops (Dinosauria, Neoceratopsia) from the Late Cretaceous of Inner Mongolia (P. R. China).
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–American Museum Novitates: #3530, pp. 1-42 - Sereno, P. C., 1990: Psittacosauria. 579-592
in Weishampel, D. B., Dodson, P., & Osmólska, H. (eds.), 1990: The Dinosauria.
–University of California Press, Berkley, Los Angeles, Oxford, 1990 xvi-733 - Sereno, P. C., 2000: The fossil record, systematics and evolution of pachycephalosaurs and ceratopsians from Asia. 480-516
in Benton, M. J., Shishkin, M. A., Unvin, D. M. & Kurochkin, E. N., (eds.) 2000: The Age of Dinosaurs in Russia and Mongolia.
–Cambridge University Press, Cambridge, xxxix-696 - Sereno, P. C. & Chao, S., 1988: Psittacosaurus xinjiangensis (Ornithischia: Ceratopsia), a new psittacosaur from the Lower Cretaceous of Northwestern China.
–Journal of Vertebrate Paleontology: Vol. 8, #4, pp. 353-365 - You, H., 2000: A new phylogeny of basal Neocetopsia.
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