After Brown, Henderson, Vinther, Fletcher, Sistiaga, Herrera & Summons, 2017
<==o †Nodosauridae Marsh, 1890a [incl. Polacanthidae Wieland, 1911] (nodosaurs; nodulipanssariselät) |--+-- †Sauroplites scutiger | |-- †Mymoorapelta maysi Kirkland & Carpenter, 1994 | `-- †Dongyangopelta yangyanensis `--+-- †Gastonia burgei Kirkland, 1998 `--+-- †Gargoyleosaurus parkpinorum Carpenter, Miles, & Cloward, 1998 [Gargoyleosaurus parkpini Carpenter, Miles, & Cloward, 1998] `--+--o †Polacanthus Owen, 1865 non Huxley, 1867 | |-- †P. foxii Owen, 1865 non Hulke, 1881 | `?- †P. rudgwickensis Blows, 1996 `--+-- †Peloroplites cedrimontanus `--+-- †Sauropelta edwardsorum Ostrom, 1970a [emend. ????] [Sauropelta edwardsi Ostrom, 1970a] `--+-- †Nodosaurus textilis Marsh, 1889b [Stegopelta landerensis Williston, 1905] `--+-- †Niobrarasaurus coleii (Mehl, 1936) Carpenter, Dilkes & Weishampel, 1995 [Hierosaurus coleii Mehl, 1936] `--+-- †Tatankacephalus cooneyorum `--+-- †Silvisaurus condrayi Eaton, 1960 `--+-- †Amanantarx ramaljonesi Carpenter et al., 1999 |--+-- †Hungarosaurus tormai Ősi, 2005 | `--+-- †Europelta carbonensis | `--+-- †Borealopelta markmitchelli Brown, Henderson, Vinther, Fletcher, Sistiaga, Herrera & Summons, 2017 | `-- †Pawpawsaurus campbelli Lee, 1996 `--+--+-- †Stegopelta landerensis Williston, 1905 | `--o †Struthiosaurus Bunzel, 1871 [taxonomic history] | |-- †S. austriacus Bunzel, 1871 [taxonomic history] | |-- †S. transylvanicus Nopcsa, 1915 [taxonomic history] | `-- †S. languedocensis Carcia & Pereda Suberbiola, 2003 [taxonomic history] `--+-- †Panoplosaurus mirus Lambe, 1919 `==o †Edmontonia Sternberg, 1928 [ Chassternbergia Bakker, 1988; Denversaurus Bakker, 1988] |-- †E. longiceps Sternberg, 1928 |-- †E. rugosidens (Gilmore, 1930) [Paleoscincus rugosidens Gilmore, 1930; Chassternbergia rugosidens (Gilmore, 1930) Bakker, 1988] `-- †Denversaurus schlessmani Bakker, 1988 [Edmontonia schlessmani (Bakker, 1988)]
Polacanthines?
†Hylaeosaurus armatus Mantell, 1833 [Synonyms] †Hoplitosaurus marshi (Lucas, 1901) Lucas, 1902a [Stegosaurus marshi Lucas, 1901; Polacanthus marshi (Lucas, 1901)]
†Nodosauridae incertae sedis:
†Anoplosaurus curtonotus Seeley, 1879 †Propanoplosaurus marylandicus Stanford, Weishampel & Deleon, 2011 †Texasetes pleurohalio Coombs, 1995 †Priconodon crassus Marsh, 1888 †Liaoningosaurus paradoxus Xu, Wang & You, 2001 †Dracopelta zbyszewskii Galton, 1980a †Hierosaurus sternbergii Wieland, 1909 †Sarcolestes leedsi Lydekker, 1893c †Edmontonia australis Ford, 2000 †Nodosaurus ischyrus (Seeley, 1881)
Alternative phylogenies of Nodosauridae:
After W. L. Parsons & K. M. Parsons, 2009
<==o Nodosauridae
|-- †Sauropelta
|-- †Silvisaurus
`--+-- †Pawpawsaurus
`--+-- †Panoplosaurus
`-- †Edmontonia
|
After Carpenter, 2001
<==o †Nodosauridae
|?- †Propanoplosaurus marylandicus
|--+-- †Panoplosaurus mirus
| `--+-- †Stegopelta landerensis
| `--+-- †Niobrarasaurus coleii
| `--+-- †Texasetes pleurohalio
| `--+-- †Amanantarx ramaljonesi
| `--o †Edmontonia
| |-- †E. longiceps
| `-- †E. rugosidens
`--+-- †Anoplosaurus curtonotus
`--+--+-- †Nodosaurus textilis
| `--o †Struthiosaurus
| |-- †S. austriacus
| |-- †S. transylvanicus
| `-- †S. languedocensis
`--+-- †Sauropelta edwardsorum
`--+-- †Pawpawsaurus campbelli
`-- †Silvisaurus condrayi
|
Reference(s):
- Brown, C. M., Henderson, D. M., Vinther, J., Fletcher, I., Sistiaga, A., Herrera, J. & Summons, R. E., 2017: An Exceptionally Preserved Three-Dimensional Armored Dinosaur Reveals Insights into Coloration and Cretaceous Predator-Prey Dynamics.
–Current Biology: Vol. 27, #16, pp. 2514-2521 [doi: 10.1016/j.cub.2017.06.071] - Carcia, G. & Pereda Suberbiola, X., 2003: A new species of Struthiosaurus (Dinosauria: Ankylosauria) from the Upper Cretaceous of Villeveyrac (southern France).
–Journal of Vertebrate Paleontology: Vol. 23, #1, pp. 156-165 - Carpenter, K., 2001: Phylogenetic analysis of the Ankylosauria. xv-526
in Carpenter, K., (ed.) 2001: The Armored Dinosaurs.
–Indiana University Press, Bloomington & Indianapolis, 2001, pp. xv-526 - Carpenter, K., Dilkes, D. W. & Weishampel, D. B., 1995: The dinosaurs of the Niobrara Chalk Formation (Upper Cretaceous, Kansas).
–Journal of Vertebrate Paleontology: Vol. 15, #2, pp. 275-297 - Coombs, W. P. Jr., & Marya´nska, T., 1990: Ankylosauria. 456-483
in Weishampel, D. B., Dodson, P., & Osmólska, H. (eds.), 1990: The Dinosauria.
–University of California Press, Berkley, Los Angeles, Oxford, 1990 xvi-733 - Keesey, T. M., 1996-2000: Dinosauricon
–iNet: The Dinosauricon: http://dinosauricon.com/ - Lee, Y.-N., 1996: A new nodosaurid ankylosaur (Dinosauria: Ornithischia) from the Paw Paw Formation (Late Albian) of Texas.
–Journal of Vertebrate Paleontology: Vol. 16, #2, pp. 232-245 - Parsons, W. L. & Parsons, K. M., 2009: A new ankylosaur (Dinosauria: Ankylosauria) from the Lower Cretaceous Cloverly Formation of central Montana.
–Canadian Journal of Earth Sciencies: Vol. 46, pp. 721-738 [doi: 10.1139/E09-045] - Stanford, R., Weishampel, D. B. & Deleon, V. B., 2011: The First Hatchling Dinosaur Reported from the Eastern United States: Propanoplosaurus marylandicus (Dinosauria: Ankylosauria) from the Early Cretaceous of Maryland, U.S.A.
–Journal of Paleontology: Vol. 85, #5, pp. (916-924) [doi: 10.1666/10-113.1] - Suberbiola, X. P. & Galton, P. M., 2001: Reappraisal of the nodosaurid ankylosaur Struthiosaurus austriacus Bunzel from the Upper Cretaceous Gosau Beds of Austria. 173-210
in Carpenter, K., (ed.) 2001: The Armored Dinosaurs.
–Indiana University Press, Bloomington & Indianapolis, 2001, pp. xv-526 - Xu, X., Wang, X.-l. & You, H.-l., 2001: A juveline ankylosaur from China.
–Naturwissenschaften: Vol. 88, #6, pp. 297-300 - Ősi, A., 2005: Hungarosaurus tormai, a new ankylosaur (Dinosauria) from the Upper Cretaceous of Hungary.
–Journal of Vertebrate Paleontology: Vol. 25, #2, pp. 370-383