After Ortega, Gasparini, Buscalioni & Calvo, 2000, Turner & Calvo, 2005, Carvalho, Campos & Nobre, 2005, Company, Pereda Suberbiola, Ruiz-Omeñaca & Buscalioni, 2005, and Sereno & Larsson, 2009
Alternative phylogeny of Ziphosuchia after Tykoski, Rowe, Ketcham & M. W. Colbert, 2002
Alternative phylogeny of Ziphosuchia after Pol, 2003
<==o †Ziphosuchia sensu Ortega, Gasparini, Buscalioni & Calvo, 2000, and Turner & Calvo, 2005 |?- †Wanosuchus atresus |--o †Notosuchia [Notosuchidae sensu lato] | |?- †Adamantinasuchus navae Nobre & Carvalho, 2006; U.Cret. SA. | |?-o †Comahuesuchidae Bonaparte, 1991 [placed in Notosuchia by Sereno, Larsson, Sidor & Gado, 2001, and Sereno, Sidor, Larsson & Gado, 2003] | | |-- †Comahuesuchus brachibuccalis | | `-- †Anatosuchus minor Sereno, Sidor, Larsson & Gado, 2003 (“duck-croc”; widenosed crocodile from Niger) | |--o †Candidodontidae Carvalho, Ribeiro & Avilla, 2004 | | |--o †Mariliasuchus Carvalho & Bertini, 1999; U.Cret. SA. | | | |-- †M. amarali Carvalho & Bertini, 1999 | | | `-- †M. robustus | | |-- †Candidodon itapecuruense Carvalho & Campos, 1988; L.Cret. ESA. | | `?- †Malawisuchus mwakayungutiensis Gomani, 1997; L.Cret. EAf. | `--o †Notosuchidae Dollo, 1914 [1924] | |-- †Notosuchus terrestris Woodward, 1896; U. Cret. WSA. | `--+-- †Sphagesaurus huenei Price, 1950, L. Cret. ESA. | `-- †Chimaerasuchus paradoxus Wu, Sues & Sun, 1995; L. Cret. EAs. `--+-- †Libycosuchus Brevirostris Stromer, 1914; L:Cret. NEAf. [Libycosuchidae] `--+--o †Doratodon Seeley, 1881; U.Cret. SWEu. | |-- †D. carcharidens (Bunzel, 1871) | `-- †D. ibericus Company, Pereda Suberbiola, Ruiz-Omeñaca & Buscalioni, 2005 `--o †Sebecosuchia Colbert, 1946 [Baurusuchoidea Price, 1945 sensu Carvalho, Ribeiro & Avilla, 2004] |--o †Sebecosuchidae Simpson, 1937 [Sebecidae Simpson, 1937; incl.? Bretesuchidae] | |?- †Ayllusuchus fernandezi Gasparini, 1984; Eoc. SAm. | |?- †Barinasuchus arveloi Paolillo & Linares, 2007; Eoc.-Mioc. SAm. | |?- †Ilchunaia parca Rusconi, 1946; M.Eoc. SAm. | |?- †Langstonia huilensis (Langston, 1965) Paolillo & Linares, 2007 [Sebecus huilensis Langston, 1965] Mioc. SAm. | |-- †Sebecus icaeorhinus Simpson, 1937; Mioc. SAm. | `--o †Bretesuchidae Gasparini, Fernández & Powell, 1993 sensu Paolillo & Linares, 2007 | |-- †“Itaborai crocodyliform” | |-- †Bretesuchus bonapartei Gasparini, Fernández & Powell, 1993 | `-- †Zulmasuchus querejazus (Buffetaut & Marshall, 1991) Paolillo & Linares, 2007 [Sebecus querejazus Buffetaut & Marshall, 1991; Sebecus carajazus Gasparini, Fernández & Powell, 1993 lapsus calami] `--o †Baurusuchidae Price, 1945 sensu Carvalho, Ribeiro & Avilla, 2004, and Carvalho, Campos & Nobre, 2005 |-- †Pehuenchesuchus enderi Turner & Calvo, 2005 |--o †Iberosuchus Antunes, 1975; Eoc. WEu. [Iberosuchidae] | |-- †I. macrodon Antunes, 1975 | `?- cf. †Iberosuchus Ortega et al., 1996 [Atacisaurus crassiproratus Astre, 1931] |-- †Eremosuchus elkoholicus Buffetaut, 1989; Eoc. NAf. `--o †Baurusuchidae Price, 1945 |?- †Bergisuchus dietrichbergi (Berg, 1966) Eoc. CEu. [is a trematochampsid (Buffetaut, 1988)] |?- †Campinasuchus dinizi Carvalho, et al., 2011; U.Cret. SAm. |?- †Wargosuchus australis martinelli & Pais, 2008; U.Cret. SAm. |--o †Baurusuchus Price, 1945; U.Cret. SAm. | |-- †B. pachecoi Price, 1945 | |-- †B. salgadoensis Carvalho, Campos & Nobre, 2005 | `-- †B. albertoi Nascimento & Zaher, 2010 |-- †Cynodontosuchus rothi Woodward, 1896; U.Cret. SAm. [=? Baurusuchus Gasparini, 1996, Carvalho, Campos & Nobre, 2005] |-- †Pabwehshi pakistanensis Wilson, Malkani & Gingerich, 2001; U.Cret. SAs. `-- †Stratiotosuchus maxhechti Campos, Suarez, Riff, Kellner, 2001; U.Cret. SAm.
Alternative phylogenies of Ziphosuchia:
After Tykoski, Rowe, Ketcham & M. W. Colbert, 2002<==o †Ziphosuchia |--o †Comahuesuchidae | |-- †Comahuesuchus | `-- †Anatosuchus `--+--+--o †Libycosuchidae | | `-- †Libycosuchus | `--o †Notosuchia | `--o †Notosuchidae | |--+-- †Notosuchus | | `--+-- †Chimaerasuchus | | `-- †Sphagesaurus | `--+--+?- †Candidodon | | `-- †Malawisuchus | `--o †Uruguaysuchidae | `--o †Uruguaysuchus | |-- †U. aznarezi | `-- †U. terrai `--o †Sebecosuchia |--o †Baurusuchidae | |?- †Bergisuchus | |-- †Baurusuchus | |-- †Cynodontosuchus | `-- †Pabwehshi `--+--o †Iberosuchidae | `-- †Iberosuchus `--o †Sebecosuchidae |?- †Ilchunaia |?- †Ayllusuchus |--+-- †“Itaborai crocodyliform” | `-- †Bretesuchus `-- †Sebecus |
After Pol, 2003, and Zaher, Pol, A.B. Carvalho, Riccomini, D. Campos, & Nava, 2006<==o †Ziphosuchia |--o †Uruguaysuchidae | `-- †Uruguaysuchus `--+-- †Simosuchus `--+--+-- †Malawisuchus | `-- †Candidodon `--+--o †Notosuchidae | `-- †Notosuchus `--+--o †Comahuesuchidae | |?- †Anatosuchus | `-- †Comahuesuchus `--+--+-- †Chimaerasuchus | `-- †Sphagesaurus `--o †Sebecosuchia |--o †Baurusuchidae | |?- †Bergisuchus | |-- †Stratiotosuchus | |-- †Baurusuchus | |-- †Cynodontosuchus | `-- †Pabwehshi `--+--o †Iberosuchidae | `-- †Iberosuchus `--o †Sebecosuchidae |?- †Ilchunaia |?- †Ayllusuchus |--+-- †“Itaborai crocodyliform” | `-- †Bretesuchus `-- †Sebecus Simpson, 1937 |
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