Crocodylomorpha

After Clark, Sues & Berman, 2000 and Sues, Olsen, Carter & Scott, 2003

Family-level taxonomy of all crocodylomorphs [Crocodylia sensu lato]

Alternative phylogeny of basal crocodylomorphs after Parrish, 1993, with paraphyletic “Sphenosuchia”

Alternative phylogeny of basal crocodylomorphs after Clark, Xu, Forster & Wang, 2004, with paraphyletic “Sphenosuchia”

Alternative phylogeny of monophyletic Sphenosuchia after Wu & Chatterjee, 1993

<==o Crocodylomorpha Hay, 1930 sensu Walker, 1970 [Crocodylia sensu lato] (crocodile-like archosaurs; krokotiilimaiset arkosaurit)
   |?- †Barbarenasuchus brasiliensis Mattar, 1987
   |?- †Clarencea gracilis Brick, 1959
   |?- †Dyoplax arenaceus
   |?- †Hallopus victor (Marsh, 1877)
   |?- †Macelognathus valens Marsh, 1884
   |?- †Parrishia mccreai Long & Murry, 1995 [n.d.]
   |?- †Trialestes romeri (Reig, 1963) [Trialestia: Trialestidae] [probably a sphenosuchian]
   `--+--o Sphenosuchia Haughton, 1924 sensu Clark, Sues & Berman, 2000 (sphenosuchians, whippet-crocodiles; sfenosukkit, juoksijakrokotiilit)
      |  |?- †Litargosuchus leptorynchus Clark & Sues, 2002
      |  |?- †Redondavenator quayensis Nesbitt, Irmis, Lucas & Hunt, 2005
      |  |--o †“Terrestrisuchidae” [sensu Sereno & Wild, 1992]
      |  |  |?- †Pedeticosaurus leviseuri Van Hoepen, 1915 [n.d.? ()]
      |  |  |-- †Litargosuchus leptorynchus Clark & Sues, 2002 [Pedeticosaurus sp.]
      |  |  `-- †Terrestrisuchus gracilis Crush, 1984
      |  `--+-- †Pseudhesperosuchus jachaleri Bonaparte, 1969
      |     `--o ?†Sphenosuchidae Haughton, 1924 sensu Clark, Sues & Berman, 2000
      |        |-- †Saltoposuchus connectens Huene, 1921 [S. longipes]
      |        |--+-- †Hesperosuchus agilis Colbert, 1952
      |        |  |-- †Dromicosuchus grallator Sues, Olsen, Carter & Scott, 2003
      |        |  `-- †Kayentasuchus walkeri Clark & Sues, 2002
      |        `--+-- †Sphenosuchus acutus Haughton, 1915 [Sphenosuchidae]
      |           |-- †Dibothrosuchus elaphros Simmons, 1965 [incl. D. xingsuenensis Wu, 1986]
      |           `?- †Junggarsuchus sloani Clark, Xu, Forster & Wang, 2004
      `-- Crocodyliformes

Note:

Sphenosuchia is a problematic taxon. It's monophyly is doubtful and it has been thrown from a monophyletic to paraphyletic several times.

Reference(s):

Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698.
Carroll, R. L., 1988: Appendix. 594-648.
in Carroll, R. L. 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698.
Clark, J. M. 1994: Patterns of evolution in Mesozoic Crocodyliformes.84-97.
in Fraser, N. C. & Sues, H-D. 1994: In the Shadow of the Dinosaurs.
–Cambridge University Press, New York, 1994, x-435
Clark, J. M. & Sues, H.-D., 2002: Two new basal crocodylomorph archosaurs from the Lower Jurassic and the monophyly of Sphenosuchia.
–Zoological Journal of the Linnean Society: Vol. 136, #1, pp. 77-95
Clark, J. M., Sues, H.-D. & Berman, D. S., 2000: A new specimen of Hesperosuchus agilis from the Upper Triassic of New Mexico and the interrelationships of basal crocodylomorph archosaurs.
–Journal of Vertebrate Paleontology: vol. 20, #4, pp. 683-704
Clark, J. M., Xu, X., Forster, C. A. & Wang, Y., 2004: A Middle Jurassic ‘sphenosuchian’ from China and the origin of the crocodylian skull.
–Nature: Vol. 430, pp. 1021-1024
Göhlich, U. B., Chiappe, L. M., Clark, J. A. & Sues, H.-D., 2005: The systematic position of the Late Jurassic alleged dinosaur Macelognathus (Crocodylomorpha: Sphenosuchia).
–Canadian Journal of Earth Sciencies: Vol. 42, pp. 307-321
Parrish, J. M. 1993: Phylogeny of the Crocodylotarsi, with reference to archosaurian and crurotarsan monophyly.
–Journal of Vertebrate Paleontology: vol. 13, #3, pp. 287-308
Parrish, J. M. 1991: A new specimen of an early crocodylomorph (cf. Sphenosuchus sp.) from the Upper Triassic Chinle Formation of Petrified Forest National Park, Arizona.
–Journal of Vertebrate Paleontology: vol. 11, #2, pp. 198-212
Sereno, P. C. 1991: Basal Archosaurs: phylogenetic relationships and functional implications.
–Journal of Vertebrate Paleontology: Vol. 11, Supplement to #4, Memoir 2. pp. ii-53
Sereno, P. C. & Wild, R., 1992: Procompsognathus: theropod, "thecodont" or both?
–Journal of Vertebrate Paleontology: Vol. 12, #4, pp. 435-458
Sues, H.-D., Clark, J. M. & Jenkins Jr, F. A., 1994: A review of the Early Jurassic tetrapods from the Glen Canyon Group of the American Southwest. pp. 284-294
in Fraser, N. C. & Sues, H.-D. (eds.), 1994: In the Shadow of the Dinosaurs - Early Mesozoic Tetrapods.
–Cambridge University Press, Cambridge, New York, x-435
Sues, H.-D., Olsen, P. E., Carter, J. G. & Scott, D. M., 2003: A new crocodylomorph archosaur from the Upper Triassic of North Carolina.
–Journal of Vertebrate Paleontology: Vol. 23, #2, pp. 329-343
Wu, X.-C. & Chatterjee, S., 1993: Dibothrosuchus elaphros, a crocodylomorph from the Lower Jurassic of China and the phylogeny of the Sphenosuchia.
–Journal of Vertebrate Paleontology: Vol. 13, #1, pp. 58-89

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