After Frost, 2000-2002 [AMNH Amphibian species of the World]
<==o Hyperoliinae ()? [Frost: 13/167]
|-- Acanthixalus spinosus
|--o Afrixalus sensu Frost, 2000
| |-- Kassinula wittei Laurent, 1940 [Afrixalus wittei (Laurent, 1940)]
| `-- Afrixalus ()? [29]
|-- Alexteroon obstetricans
|-- Arlequinus krebsi
|-- Callixalus pictus
|-- Chlorolius koehleri
|-- Chrysobatrachus cupreonitens
|--o Cryptothylax ()? [2]
| |-- Cryptothylax greshoffii
| `-- Cryptothylax minutus
|--o Heterixalus ()? [9]
| |-- H. alboguttatus
| |-- H. andrakata
| |-- H. betsileo
| |-- H. boettgeri
| |-- H. luteostriatus
| |-- H. madagascariensis
| |-- H. punctatus
| |-- H. rutenbergi
| `-- H. variabilis
|-- Hyperolius (ruokosammakot) [116]
|-- Kassinula wittei
`--o Nesionixalus
|-- N. molleri
`-- N. thomensis
Reference(s):
Carroll, R. L., 1988: Vertebrate paleontology and evolution.
--W. H. Freeman and company, New York, 1988, 698.
Carroll, R. L., 1988: Appendix. 594-648.
in Carroll, R. L. 1988: Vertebrate paleontology and evolution.
--W. H. Freeman and company, New York, 1988, 698.
Frost, D. R., 2000: Amphibian species of the World: An Online Reference.
–iNet: American Museum of Natural History, Department of Herpetology: http://research.amnh.org/herpetology/amphibia/index.html